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HomeMy WebLinkAbout; Ecological Resources of Agua Hedionda Lagoon; Ecological Resources of Agua Hedionda Lagoon; 1995-09-011994 and 1995 Field Survey Report of the Ecological Resources of Agua Hedionda Lagoon Submitted To: San Diego Gas and EJectric Company 4600 Carlsbad Boulevard Carlsbad, CA 92008 Submitted By; MEC Analytical Systems, Inc. 2433 Impala Drive Carlsbad, California 92008 September 1995 AGUA HEDIONDA LAGOON September 1995 TABLE OF CONTENTS 1.0 INTRODUCTION 1 1.1 Environmental Setting 1 1.2 Dredging History 3 2.0 METHODS 7 2.1 Eelgrass Reconnaissance Survey 7 2.2 Shoreline Habitat Reconnaissance Survey 8 2.3 Bird Surveys 8 2.4 Fish Surveys 11 2.5 Benthic Surveys 13 3.0 RESULTS AND DISCUSSION 16 3.1 Sensitive Habitats 16 3.1.1 Distribution of Eelgrass 16 3.1.2 Distribution of Salt Marsh 22 3.1.3 Historical Comparison 23 3.2 Birds 25 3.2.1 Species Composition, Density, and Distribution 25 3.2.2 Human Use and Disturbance 30 3.2.3 Special Status Species 31 3.2.4 Historical Comparison 32 3.3 Fish 33 3.3.1 Species Composition, Density, and Distribution 33 3.3.2 Sensitive Species 36 3.3.3 Historical Comparison 36 3.4 Benthic Invertebrates 37 3.4.1 Species Composition, Density, and Distribution of Macroinvertebrates \. 38 3.4.2 Species Composition, Density, and Distribution of Infauna .... 40 3.4.3 Sensitive Species 42 3.4.4 Historical Comparison 42 4.0 SUMMARY 44 LITERATURE CITED 46 APPENDIX A BIRD DATA APPENDIX B FISH DATA APPENDIX C BENTHIC DATA MEC Analytical Systems, Inc. AGUA HEDIONDA LAGOON September 1995 1.0 INTRODUCTION This report summarizes the results of field surveys conducted between April 1994 and June 1995 at Agua Hedionda Lagoon. The purpose of these surveys was to characterize the ecological resources in the aquatic and shoreline habitats of the lagoon. The survey designs and methods were developed in consultation with the California Department of Fish and Game, National Marine Fisheries Service, U.S. Army Corps of Engineers, and U.S. Fish and Wildlife Service during an interagency meeting on June 24, 1994. Survey plans were submitted to and approved by the agencies prior to initiation of the field work. At the time of the survey planning, a dredging project for the middle and inner sections of the lagoon was being considered by San Diego Gas and Electric Company (SDG&E), which operates a power plant adjacent to the lagoon. The surveys were designed to provide adequate environmental information to support agency review of a dredging project. 1.1 Environmental Setting Agua Hedionda Lagoon is a "man-made" water body located on the southern California coastline within the incorporated area of the City of Carlsbad (Figure 1.1). It extends 1.7 miles inland, and is up to 0.5 mile wide (Bradshaw et al. 1976). The lagoon is transected by three major thoroughfares: Carlsbad Boulevard (Highway 101) at the coastline; Atchison Topeka and Santa Fe (AT&SF) railroad, about 1500 ft to the east; and Interstate 5 (I-5), about 1000 ft farther east. The roadbeds of these thoroughfares effectively divide the water body into three sections, commonly referred to as the outer, middle, and inner lagoons. Acreage encompassed within the high tide mark (i.e., the subtidal and intertidal areas) is about 66 acres in the outer lagoon, 23 acres in the middle lagoon, and 197 acres in the inner lagoon (P. Pike, SDG&E, personal communication, 1994). The western shore of the outer lagoon is bounded by Carlsbad Boulevard. The southern shore of the outer and middle lagoons is occupied by SDG&E's Encina power plant and related support facilities. The southern shore of the inner lagoon is undeveloped with adjacent land being used for agriculture. Development is adjacent to most of the northern shore of the lagoon except for some marshy and unvegetated flats at the eastern end of the inner lagoon. The eastern extreme of the inner lagoon is undeveloped. The lagoon was constructed in 1954 to provide cooling water for the Encina Power Plant. Waters enter the lagoon from the ocean through an inlet channel located at the northwestern end of the outer lagoon. The power plant draws waters from the outer lagoon, and discharges the cooling water into a separate discharge channel that empties directly into the ocean. MEC Analytical Systems, Inc. AGUA HEDIONDA LAGOON September 1995 Anaheim Bay Upper Newport Bay ORANGE COUNTY San Clemente Pacific Ocean San Mateo Marsh Las Floras Marsh Santa Margarita River Estuary San Luis Rey River Marsh SAN DIEGO COUNTY Oceanside Carlsbad PtcilicOettn RIVERSIDE COUNTY Buena Vista Lagoon AGUA HEDIONDA LAGOON'' Batiquitos Lagoon San Elijo Lagoon )^> So/ana Beach San Dieguito Lagoon Del Mar GALJFOm*IA KCV MAP Las Penasquitos Lagoon Mission Bay San Diego Bay Tijuana Estuary San Diego MEXICO SOUTHERN CALIFORNIA KEY MAR Figure 1.1. Geographic map showing the location of Agua Hedionda Lagoon. MEC Analytical Systems, Inc. AGUA HEDIONDA LAGOON September 1995 Tides in the lagoon are semi-diurnal, ebbing and flooding twice each day. As is typical for the area, flushing rates vary as a result of bimonthly tidal range fluctuations (i.e., spring and neap tides). Tide elevations fluctuate between extremes of -4.6 and +4.3 ft National Geodetic Vertical Datum (NGVD) (Jenkins and Wasyl 1993). Agua Hedionda Creek and its main tributary, Buena Creek, drain a 29 square mile area from the San Marcos Mountains to Agua Hedionda Lagoon (USACOE 1973). The stream channel and flood plain vary along its course; the channel is shallow and indistinct with a broad flood plain adjacent to the lagoon. Mean seasonal precipitation for the drainage area ranges from about 11 inches near the coast to about 18 inches in the mountains with most rainfall between December and March (USACOE 1973; Wells and Price 1989). Freshwater enters the lagoon via Agua Hedionda Creek during the rainy season, and also enters via storm drains, urban runoff along the northern shore, and agricultural runoff along the southern shore (Bradshaw et al. 1976; Wells and Price1989). Twenty-five major storm drains have outfalls that discharge into Agua Hedionda Lagoon; no illicit connections or illegal discharges were identified in the Agua Hedionda watershed during dry weather storm water monitoring in 1993 (N. Diele, City of Carlsbad, personal communication, 1994). In addition to the power plant, the lagoon supports several other uses. SDG&E provides a parking lot for public access and fishing (Encina Fishing Area) along the western shore of the outer lagoon. Mariculture of mussels and oysters is conducted by a private firm in the outer lagoon. A YMCA camp is on the western shore of the middle lagoon, and the area is used for boating (non-power) and fishing. The inner lagoon is used for recreational water sports, particularly boating, water skiing, and jet skiing (Figure 1.2). 1.2 Dredging History SDG&E purchased the Agua Hedionda slough and adjacent property along the southern shore in the 1950's to construct the Encina Power Plant and a tidal basin to provide its cooling water. The slough was a silt choked, 230 acre marsh that only occasionally was open to the sea (Lillevang 1966; Bradshaw et al. 1976). Presumably, odors resulting from stagnant water conditions is how the slough got its Spanish name, which means stinking water. SDG&E and private contractors dredged about 4.3 million cubic yards (cu yd) of sediment from the slough in 1954, attaining depths of about 8 ft below mean sea level (MSL) (7.8 ft NGVD) throughout most of the lagoon (Ellis 1954). Two settling basins were included in the design: one in the outer lagoon (-20 ft MSL) to trap sands carried in from the ocean, and one at the eastern end of the inner lagoon (-17 ft MSL) to trap incoming sediments from the Agua Hedionda Creek (Leeds 1953; Jenkins and Skelly 1988). MEC Analytical Systems, Inc. II II II fill II I I II II II II II II II II II II fill S^. 01 f S' I I.« CQ I ro I O Q' o CQ vo {aO Q. CO CO a6' CO CD 0)T3 CD 5' CD CO CDa6' CQ V SNUG HARBOR MARINA (Privately Owned)Public Launch Ramp Watercraft Area &® & Use Areas - Year Round BRISTOL COVE PrivateResidents Only ^ X CARLSBAD BOAT CLUB Private^ Members Only 7^\ x. • ^\ Powerboat Area No Passive Vessels \^ ^N>^*- /\^ </x ^w A A %^/-•*£*> . SW Beach SE * Existing Buoys ^^ A Proposed Buoys Mil/ A X v\\\ \ \ } } \ Public Access / / \ Wa|k-|n-°n|y / / ^ — tt ^^^C ///* \ ///\ p/ ^-^. * \ X./ Passive Use Area ' :'?N^ / Boardsail & Other .• •.•!::5i-'.-?Nv^ Approved Vessels V. •,''.\-V:i'v-Vi'??Xw Existing ''•'^•'-''\'^-!:i'^>^* Slalom Course " •"•>••'/ H A ••'.• ••.''•'•'.'•- ''::.\'':'-d-i''":* ~*!**8$\ P^posed"" ' ..vJ'^'-j^ Beach '\ Slalom Course tjs^ OGtk 5 1S r» Ss Oi $0 S1 i 1 *•*, AGUA HEDIONDA LAGOON September 1995 The lagoon functions as an effective coastal sediment trap due to its short and porous jetties and rectification of tidal currents (i.e., ebbing currents weaker than flooding currents) due to the diversion of lagoon water through the power plant condensers (Jenkins and Skelly 1988). Maintenance of the lagoon since its construction has been limited to the dredging of littoral sands that become trapped in the outer lagoon, and dredging has taken place every one to three years since 1955 (Jenkins and Skelly 1988; USACOE 1993). Dredged materials have been used to replenish the beach both north and south of the lagoon inlet. No dredging has occurred in the middle and inner lagoons since construction. Substantial sediment infilling has occurred in the outer lagoon since the 1993 dredge cycle, and in the middle and inner lagoons since their construction. Large sandbars occur in the outer and middle lagoons, and a pair of sandbars are found at the western end of the inner lagoon (Figure 1.3). The formation of sandbars in the middle and inner lagoons was first noticed in 1960 (Bitter 1972), and they have greatly expanded since then. Grain size analyses indicate that sandy sediments, which result from the intrusion of beach sands, characterize the middle lagoon deposits (MEC 1993a). The bars in the west inner lagoon are sandy also, but with a higher silt/clay content than the bar in the middle lagoon. The -17 ft. MSL sediment trap in the inner lagoon has filled entirely. A sediment delta of fine sediments has developed at the eastern end, extending 1000 ft or more into the lagoon. Runofff from surrounding agricultural fields, urban development construction, and the Agua Hedionda Creek watershed are considered sources of these sediments (Leighton and Associates 1988 cited in Jenkins and Skelly 1988; MEC 1993a). «• MEC Analytical Systems, Inc. to AGUA HEDIONDA LAGOON September 1995 Inlet Channel PACIFIC OCEAN Discharge fc^ Channel 1009 FEET 700 «OOUCTEKS •'""."."_> Sand bar or delta Figure 1.3. Locations of sand bars and the sediment delta in Agua Hedionda Lagoon. MEC Analytical Systems, Inc. AGUA HEDIONDA LAGOON September 1995 2.0 METHODS Field reconnaissance surveys of eelgrass and sensitive shoreline habitats were performed in April 1994. Monthly bird surveys were conducted from June 1994 through June 1995. Fish and macroinvertebrates were surveyed during the summer (July 1994) and spring (April 1995) seasons when highest abundances are expected. Benthic infaunal invertebrates also were surveyed during the spring fish survey. The survey plans for each of these resource components are described in the following subsections. On tables and in the text, distances are reported in feet (ft) and/or meters (m). Mean sea level (MSL) at Agua Hedionda Lagoon is +0.19 ft NGVD (Jenkins and Wasyl 1993); mean lower low water (MLLW) is equivalent to -2.8 ft MSL (or -2.6 ft NGVD). Water depths at the time of sampling are reported for fish and infauna sampling. 2.1 Eelgrass Reconnaissance Survey A reconnaissance survey to determine the distribution of eelgrass in the outer, middle, and inner lagoons was conducted April 18-20,1994 during neap tide conditions. The location of eelgrass was determined by SCUBA divers in the outer and middle lagoons. SCUBA diving in the inner lagoon proved ineffective due to poor visibility. Eelgrass in the shallow inner lagoon was more efficiently observed from the water surface and periodically confirmed by diving. Divers noted whether the eelgrass was patchy or continuous in distribution, and took one to three density measurements (number of stems per 0.25 m2) in representative bed areas. The distribution of eelgrass was mapped by marking the lagoon^ward edges of the eelgrass beds with weighted pop-float buoys. The latitude and longitude of the pop-float buoys were located with a Magnavox X200 Global Positioning System (GPS). Eelgrass locations were plotted by SDG&E survey staff on bathymetric maps of the lagoon from the GPS marks. Multiple GPS readings were taken at known reference locations within each section of the lagoon, both before and after surveying eelgrass locations, to facilitate establishment of a plotting grid for mapping. The GPS system was field demonstrated to be accurate within a range of 6.5 ft (2 m) for most areas of the lagoon; accuracy was poor along the southern side of the inner lagoon probably due to signal obstruction from the high elevation bank. Map locations along the south side of the inner lagoon were adjusted based on field notes and depth contours. Eelgrass locations in the outer lagoon additionally were surveyed by SDG&E personnel using a bearing-bearing intersection technique. This technique provides accuracy within a range of 0.2 ft of the object, and is routinely used by SDG&E to accurately map eelgrass in the outer lagoon before and after maintenance dredging (P. Pike, SDG&E, personal communication, 1994). The greater degree of mapping accuracy for the outer lagoon was necessary since the field survey also was used to produce the post-dredging eelgrass map for the 1993 dredging cycle. MEC Analytical Systems, Inc. AGUA HEDIONDA LAGOON September 1995 2.2 Shoreline Habitat Reconnaissance Survey The perimeter of the middle and inner lagoons was surveyed on April 27-28, 1994 to determine the occurrence and locations of sensitive habitat such as salt marsh. Locations were determined with the aid of an aerial photograph and landmarks. The perimeter was defined as the area from the water's edge to 25 ft inland from the high tide mark. The locations of salt marsh habitat along the shore of the lagoon were added to the maps of eelgrass distribution. In addition, the 1994 reconnaissance was used to provide recent field validation of a 1993 habitat mapping, which was based on the analysis of a 1:500 scale color aerial photograph taken in March 1993 (MEG 1993b). The aerial photograph originally was interpreted by Wetlands Research Associates (WRA) using U.S. Geologic survey (USGS) topographic maps, National Wetlands Inventory (NWI) maps, and experience from previous site visits. Habitats at or below the 10-foot topographic contour line (NGVD) were delineated. The field validation focused on shoreline habitats. 2.3 Bird Surveys The purpose of the surveys was to characterize annual bird utilization of Agua Hedionda Lagoon. Reconnaissance surveys of the east inner lagoon were conducted in June and July 1994. Methods established during the reconnaissance surveys were followed during subsequent surveys of the entire lagoon. Birds were censused once each month in the entire lagoon from August 1994 through June 1995. Surveys generally were conducted in the morning hours between 8 a.m. and noon; a couple of surveys were conducted in late afternoon. Shorebird habitat was surveyed within 2 hours of low tide conditions. Censusing took place from land vantage points, and from an inflatable boat. Total saturation coverage methodology was employed, whereby all birds were counted according to species, lagoon section, and habitat. Specifics regarding census information, sampling locations, and habitat are given below. Census Information All birds were identified and counted by species. Evidence of nesting (e.g., nests, chicks, and fledglings) was separately recorded. Lagoon Sections, Habitats, and Shorelines Birds were censused in four sections of the lagoon (Figure 2.1), as follows: (1) outer lagoon (2) middle lagoon (3) west inner lagoon (i.e., west of Bristol Cove) (4) east inner lagoon (i.e., east of Bristol Cove) MEC Analytical Systems, Inc. AGUA HEDIONDA LAGOON September 1995 Inlet Channel PACIFIC OCEAN Km FEET Sand bar or delta Figure 2.1. Lagoon sections and shoreline designations for the 1994 and 1995 bird surveys at Agua Hedionda Lagoon. MEC Analytical Systems, Inc. AGUA HEDIONDA LAGOON September 1995 The inner lagoon was divided into east and west sections because of differences in sedimentation and depth. The west inner lagoon has sandbars near the channel to the middle lagoon, and maximum depths are about -8 to -9 ft. MSL. The east inner lagoon has a shallow sediment delta at the eastern extreme, and maximum depths range from -2 ft. at the eastern extreme to -8 ft. MSL near Bristol Cove (= Shelter Cove) (see Figures 3.4 and 3.5 in Section 3.0 for depth contours). Within each lagoon section, birds were censused according to the following twelve habitats: (1) Open water (2) Intertidal flat (i.e., soft substrate) (3) Bank (i.e., elevated area above high tide zone) (4) Beach (i.e., area above intertidal) (5) Rip rap (i.e., large boulders and/or rock on intertidal) (6) Sandbar or sediment delta (7) Salt marsh (8) Salt panne (9) Seasonal ponds (10) Aquaculture (=mariculture) racks (11) Other perches (e.g., docks, barges) (12) Aerial Habitats on land (i.e., bank, beach, intertidal flat, rip rap, salt marsh, salt panne, seasonal pond) data were recorded according to lagoon shoreline (e.g., east, west, north, south). For the outer lagoon, the west shoreline was defined as adjacent to Carlsbad Blvd., and the remaining shoreline was considered either north or south. The shoreline of the middle and west inner lagoon sections was recorded as either north or south. The shoreline of the east inner lagoon was recorded as either north or south (with the boundary being at the eastern extreme where the creek channel enters the lagoon) for all terrestrial habitats except for the intertidal flat. Because of the movement of birds while foraging on the intertidal flats, it was more accurate to divide the shoreline of the intertidal flat into north, south, and east shores. The north shore was defined as extending east from Bristol Cove (= Shelter Cove) to the end of the adjacent sandy beach. The south shore was defined as being on the opposite side of the lagoon from Bristol Cove to the beach on the south shore. The east shore extended from the eastern edge of the beach on the north shore to the edge of the beach on the south shore. The beach locations, as well as some of the habitats mentioned above, may be seen on Figure 3.1 in Section 3.0 of this report. The beach locations east of Bristol Cove, have a habitat code of 721 and are light in color on Figure 3.1. MEC Analytical Systems, Inc. 10 AGUA HEDIONDA LAGOON September 1995 2.4 Fish Surveys Fish were collected by beam and otter trawl nets towed by boat, and by seine nets deployed from shore. The beam trawl samples also were processed for macro- invertebrates; i.e., larger epifauna that live on the sediment surface. Surveys were conducted in July 1994 and April 1995. Water quality (dissolved oxygen, pH, salinity, temperature) and water transparency (Secchi depth) were recorded at each trawl station during the surveys. Specifics regarding sampling gear, stations, and sample analysis are given below. Sampling gear 12 ft (3.7m) otter trawl, 3.3 ft (1 m) wide beam trawl, 6 x 50 ft (1.8 x 15.2 m) beach seine with a 6 x 100 ft (1.8 x 30 m) barrier net, Hydrolab water quality analyzer, Secchi disc. Lagoon Sections and Stations Fish were surveyed in the same four lagoon sections as birds: outer lagoon, middle lagoon, west inner lagoon, and east inner lagoon. The approximate station locations for the trawl and seine sampling are shown on Figure 2.2. Depth and shoreline conditions were taken into consideration when selecting gear types for each lagoon sections. For example, seines were not used in the outer lagoon because the shoreline is lined with rip rap. The large otter trawl was not used in the middle lagoon because of eelgrass density, or in the east inner lagoon because of soft sediments and insufficient water depth. The types of samples collected in each lagoon section are summarized below. outer lagoon -1 otter trawl, 2 beam trawls middle lagoon - 3 beam trawls, 2 beach seines west inner lagoon -1 otter trawl, 2 beam trawls, 2 beach seines east inner lagoon - 2 beam trawls, 3 beach seines Equipment Deployment The otter and beam trawls were towed on the lagoon bottom from an inflatable boat for a standardized time of 5 minutes at constant traveling speed. Multiple beach seines were conducted within a barrier net set at each station. The barrier net was set with the shore serving as one edge of a 328 ft.2 (100 m2) rectangle or square. Four beach seine hauls were conducted within the barrier net, and a final (fifth) haul consisted of retrieval of the barrier net. MEC Analytical Systems, Inc. 11 AGUA HEDIONDA LAGOON September 1995 was K£T OUKETCKS Trawl Tracks July 94 and April 95 .'"" .'.".:• Sand bar or delta or April 95 Figure 2.2. Locations of beach seine (S), beam trawl (B), and otter trawl (O) stations for the 1994 and 1995 surveys of fish at Agua Hedionda Lagoon. MEC Analytical Systems, Inc.12 AGUA HEDIONDA LAGOON September 1995 Hydrolab readings were taken at the trawl stations for dissolved oxygen, pH, salinity, and temperature at the surface, mid-water, and bottom if water depths were in excess of 4 ft, or at the surface and bottom at shallower depths. The depth at which the Secchi disc was no longer visible from the water surface was recorded as a measure of water transparency. Sample Analysis All fish were counted and identified to the lowest practicable taxon (usually species). Small juvenile atherinids (< 25 mm), engraulids (< 50 mm), and gobiids (< 25 mm) were identified to family. Unless extremely abundant, fish were measured (standard length) and weighed. When there were more than 100 individuals of a species, the additional fish were measured by size class (cm) and batch weighed. Fish were field processed and released to the extent practicable. 2.5 Benthic Surveys Macroinvertebrates collected by beam trawl during the fish surveys were analyzed to provide a measure of the benthic community in the lagoon. Animals were counted and identified to the lowest practicable taxon (usually species), and released in the field to the extent possible. Small infaunal invertebrates that live within the sediment were collected in each of the same four lagoon sections surveyed for birds and fish. One survey was conducted in April 1995, to coincide with the fish survey during the spring season. Samples were collected from subtidal habitats within eelgrass beds, non-vegetated areas, on the bars that occur within the middle and inner lagoons, and on the sediment delta at the east end of the inner lagoon. Samples were collected for the analysis of grain size at half the stations. Stations A total of twenty-six stations were sampled from the different lagoon sections (Figure 2.3), and are summarized below according to lagoon section. Lagoon Section Outer Lagoon Middle Lagoon West Inner Lagoon East Inner Lagoon Total Habitat within eelgrass bed non-vegetated within eelgrass bed non-vegetated non-vegetated bar within eelgrass bed non-vegetated bar with eelgrass within eelgrass bed non-vegetated non-vegetated deltajbar) Number of benthic samples 2 2 2 2 2 2 4 2 2 2 4 26 Number grain size samples 1 1 1 1 1 1 2 1 1 1 2 13 MEC Analytical Systems, Inc.13 AGUA HEDIONDA LAGOON September 1995 "."' Sand bar or delta within edgrass bed O non-vegetated + grain size also collected Figure 2.3. Locations of stations for the 1995 survey of benthic infauna at Agua Hedionda Lagoon. MEC Analytical Systems, Inc.14 AGUA HEDIONDA LAGOON September 1995 Sampling Gear Divers used a 3 in (7.6 cm) diameter by 20 in (50 cm) deep (0.005 m2 area) hand-coring device to collect biological and sediment grain size samples. The same diameter core, but with a 6 in (15 cm) depth was used to collect samples in sandy sediments. Field Sample Processing Each biological sample was washed in the field through a 1.0 mm sieve. Material retained on the sieve was transferred to labeled quart jars, and a solution of magnesium chloride and lagoon water was added to the jar to anesthetize the organisms for about 30 minutes. Samples were then fixed with a 10% Formalin/seawater solution. Grain size samples were placed in labeled plastic bags and stored on ice for transport to the laboratory. Laboratory Analysis After at least 72 hours, the formalin solution was drained and washed from the samples, which were then preserved with 70% ethanol. Samples were sorted by major taxonomic group (annelids, crustaceans, mollusks, minor phyla, insects) by trained technicians. Taxonomists counted and identified animals to the lowest practicable taxon (usually species). Wet weight biomass was determined for each of the major taxonomic groups by transferring the vial contents to a 0.3 mm screen, vacuum-aspirating for 10 seconds, and weighing to the nearest 0.001 g using an ASP Z400 DR balance. Grain size of the sediment samples was determined for particles larger than 64 microns by gravimetric sieving, and for smaller particles by standard pipette methods (Plumb 1981). MEC Analytical Systems, Inc. 15 AGUA HEDIONDA LAGOON September 1995 3.0 RESULTS AND DISCUSSION 3.1 Sensitive Habitats Four main habitat categories occur within and adjacent to the lagoon: subtidal, flats, marsh, and upland. Several habitat maps have been prepared for the lagoon environment, the most recent by Wetland Research Associates (WRA) (MEC 1993b) (Figure 3.1). The April 1994 reconnaissance survey indicated that the previous maps are still generally valid. The focus of the 1994 reconnaissance surveys was to locate sensitive eelgrass habitat within the lagoon, and sensitive habitat within 25 ft of the shoreline. The distribution of eelgrass within the lagoon is described in Subsection 3.1.1. Intertidal flats and marsh habitats are described in Subsection 3.1.2. 3.1.1 Distribution of Eelgrass Eelgrass (Zostera marina) in the outer lagoon occurred primarily along the shoreline (Figure 3.2); its distribution is largely controlled by the agency approved limits of maintenance dredging in that section of the lagoon. Little eelgrass occurred near the inlet, and then occurred first in patches and then in larger beds along the west and northeast shores. Eelgrass was well developed along the southeast shore. Average eelgrass density ranged from 36 to 77 stems per quarter square meter. Eelgrass occurred to depths of -18 ft MSL in the outer lagoon. Eelgrass was found throughout most of the middle lagoon with the exception of the top of the sandbar, and in most of the channel between the outer and inner lagoons (Figure 3.3). Maximum depths in the middle lagoon were about - 8 to -9 ft MSL with the exception of two scour holes (depths > 20 ft MSL) in the channel between the outer and inner lagoon. Average eelgrass density in the middle lagoon ranged from 37 to 73 stems per quarter square meter, and eelgrass occurred at depths between about -2 and -9 ft MSL. Substantial eelgrass occurred on the sandbars of the west inner lagoon, and in narrow bands along the shoreline (Figure 3.4). Similar to the middle lagoon, maximum depths in the west inner lagoon were about - 8 to -9 ft MSL. However, the lower limit of eelgrass in the west inner lagoon only extended to about -4 to -5 ft MSL. Average eelgrass density along the shore ranged from 15 to 44 stems per quarter square meter, and beds started to become patchy along the south shore. Eelgrass density on the sandbar averaged 52 stems per quarter square meter. Continuing further east, eelgrass thinned to non-continuous, patchy beds with an average density of 40 stems per square meter; no eelgrass was observed at the far eastern end of the lagoon (Figure 3.5). MEC Analytical Systems, Inc. 16 Estuanne open water Coastal scrub and chaparral Southern coastal salt marsh Coastal brackish/freshwater marsh Riparian Tidal estuarine flats (sand and mud) Nontidal estuarine flats (sand and mud) Bluffs (Nonvegetated) Ruderal Agricultural Nonvegetated disturbed Urban Habitats within and adjacent to Agua Hedionda Lagoon (Source: MEC 1993b, aerial photograph interpreted by Wetlands Research Associates).* 1994 reconnaissance noted mixed habitat consisting mainly of ruderal and coastal scrub and chaparral '* 1994 reconnaissance noted some salt marsh species - ETL GRASS LDCATtt APRIL I9M • KL WASS LBCATUl JtrexXR IM3 ENCINA OUTER UGOON CARLSBAD NOTES: "OTEi AWL 1994 *VWABE KNSITY OF Ed.™*" • t Of PLANTS PERrflJKAtr •CO E - 40 STEMS10 ' - 48 ITEM JH * - » «TEMJ•JJ I* - SI STEMS«" 17 - 77 ITCMS 11/4 nrttr »a • 8.7 M jc NOTEi KCCMJER 1993 AVOMGC OCMSm CF EELDUJS • 9 PLANTS PE«XMMRE roar - AS HCASUNEIW THE THttt HAJW EELADEAi. •"on w frarm SHIET 1/4 •UN IBM a*i * mrnur BNCINA LACOONS ACUA HBDIONDA CARLSBAD . ~ PAT Figure 3.2. April 1994 distribution of eelgrass in the outer lagoon.18 I I I I I ft. IM BED e MIDDLE LAGOON BED 8-37 STEMS BED 10 - 73 STEMS NOTE: ELEVATIONS AND CONTOURS ARE JUNE 1993 SOUNDINGS Figure 3.3. April 1994 distribution of eelgrass and salt marsh in the middle lagoon. I I I I I I ENCINA INNER LAGOON BETT23 ( PATCHY EEl SRASS DENSITY AND LOCATION EC MEC ANALYTICAL SYSTEMS, INC. APRIL 199' NOTE: IN THE INNER LAROON AVERAGE DENSITY OF EEl. ORASS - | OF PLANTS PER 1/4 METER SQUARED. BCD 1« - 15 STEMS BED 20 - 40 STEMS BED 23 - 44 STEMS BED !5 « 52 STEMS BED 26 - 30 STEMS (1/4 m«t«r to. - 2.7 ft 10.' menu LAGOONS AGUA HDIONDACARLSBAD AGUA HED10NDA LAGOON AfTCK ORCDCt CIJYA110NHOTD ELEVATlOHi WW CONTOURS ARC fXOU JOHC 1»3 Figure 3.4. April 1994 distribution of eelgrass and salt marsh in the west inner lagoon.20 II I I I I ones DENStrr AND LOCATIONfl MCC ANALYTICAL 5YSTQ4S.trm.IIMER LAGOONoExsnv or EELO BRASS - I OF PIAKTS PER 1/4 METW SQUAMD. 19 - 15 BCD 20 - 40 STEMS KD 23 - 44 STEWS BED 28 » 52 STEMS BED 2« - 3C STEMS (1/4 n»tor M. - 2.7 n. -) INNER LAGOON -# I • PSDPOStl AFTER DREtCE ELEVATIDH 8i500 > VCLUHE OF THE -17 IASIH 449JM - VDLU* rmw THE -17 IAJ1N TD STA. - VDLUHE OF THE 1SLANI > TOTAL VOLUME INNER LAGOON NOTE : ELEVATIONS AND CONTOURS ARE FROM JUNE 1993 SOUNDINGS KNCTNA UGOONS AGUA HBD10ND* CARLSBAD RANCHO AGUA HED10NDA Figure 3.5. April 1994 distribution of eelgrass and salt marsh in the east inner lagoon.21 m AGUA HEDIONDA LAGOON September 1995 m 3.1.2 Distribution of Salt Marsh •• Nearly the entire shoreline of the outer lagoon is armored by rock revetment; one exception M is at the southeast shore where the intertidal is sandy with cobble patches. Rock rip rap also lines the channels between the outer and middle lagoons and between the middle and «* inner lagoons. M Salt marsh and tidal flats occur along the shores of the middle and inner lagoons. P Common salt marsh species included pickleweed (Salicomia virginica), saltgrass H (Distichilis spicata), fleshy jaumea (Jaumea camosa), and alkali heath (Frankenia grandifolia). Brackish/freshwater marsh species such as cattail (Typha latifolia), bulrush m (Scirpus sp.), and spiny rush (Juncus sp.) occur in small areas in the inner lagoon. H The middle lagoon has narrow tidal flats along each shore; the widest flats occur along the m north shore and at the eastern end of the south shore. The north shore has narrow tidal ^ flats, and pickleweed occurred above mean high water in the northwest and northeast corners, and in scattered, small patches in between (Figure 3.3). The parasitic plant Cuscuta salina was associated with the pickleweed. The east shore has a narrow bank, 11 and scattered small patches of pickleweed were scattered along this shore. «•The inner lagoon is bordered by steep banks on the north and south sides, and low lying m areas at the east end. Rocky cobble and boulder areas were noted along the northwestern m shore from the entrance channel to the Snug Harbor Marina; concrete and tire shore ta protection was being used at Snug Harbor Marina; cobble, boulders and some concrete occurred in the vicinity of Agua Hedionda Point and the Carlsbad Boat Club; and rip rap *. was observed on both sides of the entrance to Bristol Cove. Tidal flats become exposed ta during low tide along most of the shoreline of the west inner lagoon. Salt marsh plants occurred in small patches in the Snug Harbor Marina area, and extended as a narrow — fringe along the eastern shore of Snug Harbor about half-way towards Agua Hedionda ^ Point (Figure 3.4; see Figure 3.1 for locations of landmarks). Pickleweed and saltgrass were best developed just east of the Snug Harbor Marina. A small stand of cattails also — occurred in this vicinity. Banks measuring about 2 to 5 ft high occur east of the marina ». towards Agua Hedionda Point, and pickleweed occurred in scattered patches along the lower portions of the bank. Saltgrass was common on the edges of the bank. Golden "" weed (Haplopappus ericoides) and iceplant occurred on the banks. No salt marsh plants IN were observed between the Carlsbad Boat Club and the Bayshore Drive public access. Instead, non-vegetated disturbed land and ruderal (mainly weedy) vegetation were "" common. *• Mudflats were best developed at the east end of the inner lagoon, and have expanded in "" recent years due to extreme sedimentation. Sandy flats occur at the Bayshore Drive public •• access, and there are two beach areas along the southern shore of the inner lagoon that have expanded in size since the 1970's. The most extensive salt marsh occurred east of "" the Bayshore Drive public access and extended to the eastern end of the lagoon (Figure •• 3.5). This area was dominated by pickleweed, mudflat, tidal creeks, and non-tidal flats. ^ Pickleweed was best developed along the lagoon bank and tidal channels, and decreased IH MEC Analytical Systems, Inc. 22 AGUA HEDIONDA LAGOON September 1995 in vigor within a short distance from those areas where regular tidal flushing occurs. The parasitic plant Cuscuta salina occurred on some of the pickleweed. Fleshy jaumea occurred in patches near the tidal channels and along the lagoon bank. Alkali heath and saltgrass sparsely occurred in upper mid-marsh areas. 3.1.3 Historical Comparison Eelgrass The present distribution of eelgrass is less than it was twenty years ago (Figure 3.6). Twenty years ago, there was more eelgrass in the middle lagoon because of the smaller size of the sandbar. Additionally, eelgrass formed a much more continuous bed in the western section of the inner lagoon, and patches of eelgrass extended further east. Bradshaw et al. (1976) indicated that the distribution of eelgrass in Agua Hedionda Lagoon appears to be controlled by depth, substrate stability, and light availability (i.e., the water sufficiently clear for photosynthesis). Light levels were considered the primary factor controlling the density of eelgrass relative to depth in the middle lagoon by Backman and Barilotti (1976). Because of the changes that have occurred in the lagoon due to sediment infilling over the last twenty years, it is reasonable that depth, substrate stability, and light all have contributed to the present distribution of eelgrass. The decreases in eelgrass since the 1970's in the shallower eastern end of the lagoon may relate to sediment deposition, and to some infilled areas being too shallow for eelgrass. It is unknown to what extent the decrease in eelgrass in the west inner lagoon relates to turbidity or to powerboat use, either directly (e.g., by removal) or indirectly (e.g., by increased turbidity and reduced light). Eelgrass in the west inner lagoon occurred close to shore and within areas around the sandbars that are protected from boating by warning buoys (refer to Figures 1.2 and 3.4). Salt Marsh The present study indicated that the previous habitat maps are still generally valid; however, more wetland habitat was noted than previously reported. The "new" wetland areas (salt marsh) were located in the middle lagoon, and in the Snug Harbor Marina area of the inner lagoon. Previous reports of the largest expanse of salt marsh occurring at the east end of the lagoon were confirmed by the 1994 reconnaissance. The salt marsh of the east inner lagoon was characterized by Bradshaw et al. (1976) as having very limited plant diversity. This was confirmed by the 1994 reconnaissance which noted a low plant diversity and a reduced vigor of pickleweed within a short distance from the lagoon bank and tidal channels. MEC Analytical Systems, Inc. 23 0 LEGEND D I I D I I II D 0 BOUNDARY OF HABITAT STUDY AREA ESTIMATED HIGHEST JIDE OK SANDY SHORE \-\ \PELAGIC GRASS, ZOSTER A MARINA ESTIMATED LOWEST TIDE APPROXIMATE LOCATION OF CHANNELCARLSBAD VEGETATION BRACKISH WATER ALLUVIAL FAN Figure 3.6. Distribution of eelgrass and salt marsh in Agua Hedionda Lagoon in 1975 (Source: Bradshaw et al. 1976).Page 24 " AGUA HEDIONDA LAGOON September 1995 m to 3.2 Birds "" Southern California estuaries and lagoons, provide habitat for migratory as well as resident m bird populations. During the spring and fall, migrating shorebirds and waterfowl arrive to ^ refuel and rest as they make their journeys northward or southward. In addition, breeding migrants such as some terns (e.g., California Least Tern) fly specifically to southern *" California lagoons and estuaries to nest. Bird utilization at Agua Hedionda Lagoon during 1994 and 1995 is summarized in "" Subsection 3.2.1 below. Because of the many recreational uses of the lagoon, human use p. and disturbance to birds is discussed in Subsection 3.2.2. Special status species are addressed in Subsection 3.2.3., and results are compared to historical studies in Subsection 3.2.4. m Data from the field surveys are presented in Appendix A. Count data for each species by survey are presented in Appendix A.1; count data by survey, lagoon section, and habitat — are in Appendix A.2; and count data by bird category, habitat, lagoon section, and shore „. are given in Appendix A.3. "" 3.2.1 Species Composition, Density, and Distribution w A total of 81 species were identified during June 1994 through June 1995 surveys. The """ majority of the species were water associated birds, accounting for 61 species »• (approximately 75 percent of the total number of species) (Table 3.1). Of these, shorebird diversity was highest, followed by dabbling ducks/geese/coots, and diving ducks. Land "~ species, in contrast to water species, can rely on water habitats (e.g., salt marsh), but also «- utilize a wide variety of transitional and upland habitats. Land species were dominated by ^ passerines or perching birds. •" Bird abundance was highest from July 1994 through April 1995 (Table 3.2). The large ^ number of birds during this time demonstrates seasonal use of the lagoon by fall, winter, and spring migrants. The populations during these months were comprised mainly of <M waterfowl including Bufflehead, Scaup, and Ruddy Duck, as well as shorebirds such as _ Semipalmated Plover, Black-bellied Plover, and Least and Western Sandpiper (Appendix A.1). M _ The east inner lagoon had the largest number of birds during all surveys, followed by the outer, west inner, and middle lagoon sections (Table 3.3). The east inner lagoon offers an "" assortment of habitats including intertidal flat, open water, pickleweed salt marsh, salt — panne, and seasonal ponds. This combination of habitats is not found elsewhere in the ^ lagoon. The intertidal flat in the east inner lagoon was the most heavily used habitat of the entire lagoon. Shorebirds, gulls, coots, and dabbling and diving ducks comprised most of — the bird community in the east inner lagoon. On outgoing and low tides, shorebirds ^ foraged for invertebrates, and herons and egrets foraged in the shallows for invertebrates and/or fish; these birds moved to nearby salt marsh and salt panne to loaf at high tides. — In contrast, gulls, dabbling ducks, coots, and a small number of terns and skimmers used mm „, MEC Analytical Systems, Inc. 25 " AGUA HEDIONDA LAGOON September 1995 m * the exposed intertidal flat and sediment delta principally for loafing. The ducks and coots feed on vegetation and/or subtidal invertebrates, and gulls are scavengers. m The pickleweed salt marsh in the east inner lagoon provided nesting and foraging habitat for the State Endangered Belding's Savannah Sparrow, a year round resident. The salt panne/salt marsh complex also was used by a small number of nesting American Avocet "• and Black-necked Stilt (Reconnaissance Survey, Appendix A. 1). In contrast to the east inner lagoon, the outer lagoon is dominated by open water, "" aquaculture racks, rip rap, and, at extreme low tide, a sandbar. Of these habitats, the open _ water and aquaculture racks were used the most. Large numbers of pelicans, cormorants, and gulls loafed on the racks, with some gulls and a small number of diving ducks and ""* grebes foraging in the open water. The sandbar was used for loafing by gulls. ta The west inner lagoon is composed primarily of open water, with narrow bands of bank and intertidal flat, beach, rip rap, and two sandbars. The beach, rip rap, and some of the — intertidal flat, particularly that which is hard and compacted, were not valuable bird habitats. ». The open water and some small areas of intertidal flat, however, offered the most productive habitats for a variety of birds (Appendix A.3). Diving and dabbling ducks, as "" well as grebes, were commonly observed foraging in the open water. Diving ducks feed M on subtidal invertebrates and vegetation, dabbling ducks feed primarily on vegetation, and grebes eat fish and invertebrates. The sandbars had some limited use for loafing gulls and "~ shorebirds. The middle portion of the lagoon had the fewest number of birds (Tables 3.2 and 3.3). This """ section of the lagoon consists of open water, a large centrally located sandbar, rip rap, and "" a small band of intertidal flat and bank. Similar to the west inner lagoon, much of the ^ intertidal flat and bank is hard and compacted and consequently offers marginal foraging "" habitat for shorebirds. A small number of diving ducks frequented the open water, and *" some shorebirds were observed on the intertidal flat (Appendix A.3). Utilization of the _ sandbar was limited, and was used mainly by loafing gulls and shorebirds. "~ 3.2.2 Human Use and Disturbance Aqua Hedionda is a recreational lagoon. Recreational fishing, ghost shrimping, and jet and """ water skiing are permitted. Although fishing and shrimping are allowed throughout the _ lagoon, only the west inner and a small section of the east inner are open to jet and water ^ skiing. What impacts, if any, recreational use of the lagoon has on the bird community is "" difficult to assess. The jet and water skiing facility is open for only a portion of the year. — During some surveys water skiing boats were observed flushing birds out of the water and ^ off the sandbars in the west inner lagoon and off the exposed delta (bar) in the east inner lagoon. The overall effect of this or other types of recreation on the number and — distribution of birds in the.lagoon is unknown. MEC Analytical Systems, Inc. 26 AGUA HEDIONDA LAGOON September 1995 Table 3.1. Summary of total number of species and percent composition by bird category at Agua Hedionda Lagoon for monthly surveys from June 1994 through June 1995. Bird Category Number of Species Percent Composition Water Species Shorebirds Dabbling Ducks/Geese/Coots Diving Ducks Grebes Gulls Terns/Skimmers Egrets/Herons Cormorants/Pelicans Loons/Mergansers Osprey Kingfisher Total 21 9 7 5 5 4 4 2 2 1 1 61 25.9 11.1 8.6 6.2 6.2 4.9 4.9 2.5 2.5 1.2 1.2 75.2 Land Species Passerines Hawks/Vultures Total 16 6 20 19.8 4.9 24.7 MEC Analytical Systems, Inc.27 AGUA HEDIONDA LAGOON September 1995 Table 3.2. Summary of total abundance of birds by month in different sections of Agua Hedionda Lagoon for surveys from June 1994 through June 1995. Survey June 1994 July 1994 August 1 994 September 1994 October 1994 November 1994 December 1994 January 1995 February 1995 March 1995 April 1995 May 1995 June 1995 Outer Lagoon ns ns 37 80 82 1129 162 87 42 45 84 64 126 Middle Lagoon ns ns 21 7 13 1 84 118 73 28 24 6 18 West Inner Lagoon ns ns 17 12 17 251 496 223 471 224 116 25 28 East Inner Lagoon 267 2712 925 1161 983 1904 649 957 676 648 859 115 151 Total 267 2712 1000 1260 1095 3285 1391 1385 1262 945 1083 210 323 ns = not sampled MEC Analytical Systems, Inc.28 AGUA HEDIONDA LAGOON September 1995 Table 3.3. Summary of total abundance of birds by category in different sections of Agua Hedionda Lagoon for monthly surveys from August 1994 through June 1995. Bird Category Cormorants/Pelicans Kingfishers Ospreys Terns/Skimmers Egrets/Herons Loons/Mergansers Grebes Dabbling Ducks/Geese/Coots Diving Ducks Shorebirds Gulls Passerines Doves Hummingbirds HawksA/ultures Total Outer Lagoon 573 . . 22 16 4 20 8 60 31 1196 8 . . . 1938 Middle Lagoon 4 4 . 7 14 2 13 11 112 178 22 21 5 . . 393 West Inner Lagoon 20 6 3 18 28 . 121 286 441 687 237 33 . . . 1880 East Inner Lagoon 25 . 3 119 79 2 62 984 793 5552 1120 263 14 2 10 9028 MEC Analytical Systems, Inc.29 AGUA HEDIONDA LAGOON September 1995 3.2.3 Special Status Species Aqua Hedionda Lagoon is inhabited by a number of sensitive bird species, ranging from Federally endangered to California species of special concern. The following discussion focuses on endangered or threatened species encountered within the lagoon. All other sensitive species are highlighted and included in the Appendix A.2 Tables. The California Brown Pelican, listed as endangered by the federal government and the State of California, can be found along the coast of San Diego County throughout the year. At Aqua Hedionda, the largest number of Brown Pelicans consistently occurred in the outer lagoon where the species used the aquaculture racks for roosting and loafing (Appendices A.2 and A.3). The species was observed in other areas of the lagoon but in much smaller numbers and at a much lower frequency. The California Least Tern is listed as endangered by the federal government and by the State of California. California Least Terns are a fairly common, but localized breeding migrant in the County during spring and summer and are absent during fall and winter. This species does not presently nest within the lagoon, but there is a seasonally active colony at the adjacent Batiquitos Lagoon. During the surveys, a total of 15 California Least Terns were seen (Appendix A.1), with 8 observed foraging and loafing in the inner lagoon. The lagoon, does not appear to offer valuable foraging habitat to this species. The Western Snowy Plover, a small shorebird, is listed as threatened by the federal government and is a California Species of Special Concern. This species is a common migrant and winter visitor and a fairly common, but localized, breeding resident in San Diego County. Western Snowy Plovers have historically nested at Aqua Hedionda but have not in recent years. Although the east end of the lagoon appears to offer useable habitat, only six individuals were observed (in the east inner lagoon) during the study. The Belding's Savannah Sparrow is a common, but localized resident in San Diego County and is an obligate of pickleweed salt marsh. This passerine is a federal Category 2 Candidate species and is listed as endangered by the State of California. Belding's Savannah Sparrows were observed only in the east inner lagoon, where expanses of pickleweed salt marsh occur. The east inner lagoon supports a small population of this species. The number of individuals increased during the March (15 individuals), April (13 individuals), and May (14 individuals) surveys (Appendix A.1), reflecting nesting activity in this portion of the lagoon. MEC Analytical Systems, Inc. 30 m AGUA HEDIONDA LAGOON September 1995 M 3.2.4 Historical Comparison p» A complete comparison between recent survey results and historical bird information for H the lagoon is somewhat difficult. Previous records are scant and variable in coverage and little is known regarding the types of conditions when the surveys were conducted (e.g., *• high or low tide, time of day). This study is the first complete annual census of birds at the H lagoon and thus, gives a more complete and comprehensive picture of the bird community inhabiting the area. p • In general, the bird populations found at the lagoon have not changed significantly since the 1970's. A total of 61 water associated species were identified during recent surveys 1 compared to the 55 reported by Bradshaw et al. (1976) and Recon (1976). Most of the • abundant and common species such as Black-bellied Plover, Least and Western Sandpiper, Marbled Godwit, Bufflehead, Ruddy Duck, and Western Grebe observed during P past Audubon Christmas counts and Point Reyes Bird Observatory surveys have continued ** to be abundant and common to the area. |j What has changed significantly over past years is the use of the lagoon by sensitive species. In the past, the endangered California Least Tem nested on the salt panne at the p east end of the lagoon. Nine pairs nested there in 1975 (Bradshaw et al. 1976). Off-road L vehicular activities and predation led to nesting failures (Ultrasystems 1983), and the species no longer breeds in the lagoon. Similarly, the threatened Western Snowy Plover, P* once a breeding resident that was abundant in the lagoon (Bradshaw et al. 1975), is now to only sporadically observed. p- Another sensitive species, the Light-footed Clapper Rail, an endangered species, once m inhabited the lagoon but has disappeared from the area over the years. The disappearance of this species is due to disturbance, degradation and elimination of •" suitable habitat (Bradshaw etal. 1976). MEC Analytical Systems, Inc. 31 AGUA HEDIONDA LAGOON September 1995 3.3 Fish Lagoons provide important habitat for coastal marine and resident fish. An important aspect of bays and estuaries is that they may serve as nursery habitat for commercially and recreationally important coastal species such as California halibut and diamond turbot. In addition, fish in lagoons provide important forage for water-associated birds. The fish surveys of the present study were conducted during two different seasons, spring and summer. Water quality was good during both surveys, but there were differences related to season and lagoon section. Temperatures ranged from 14.8 to 16.9 °C during the spring and 20.8 to 24.8 °C in the summer. Temperatures were up to 4 °C warmer in the inner lagoon during the summer than in the outer lagoon. Salinities ranged from 23 to 32.7 ppt, with the lower values in spring due to seasonal rainfall. Visibility was only about 2 to 4 ft (0.75 to 1.25 m) during the spring, probably due to rainfall runoff. Visibility was higher in the summer with the Secchi disc being visible on the bottom throughout most of the lagoon, with the exception of the inner lagoon where visibility was limited to about 4 ft (1.25m). Water quality data from the fish surveys are presented in Appendix B.1. Fish data also are given in Appendix B. Density data are in Appendix B.2, total counts are presented in Appendix B.3, and fish size data are given in Appendix B.4. 3.3.1 Species Composition, Density, and Distribution A total of twenty-nine species of fish were found during the July and April surveys of Agua Hedionda Lagoon (Table 3.4). Fewer taxa occurred in the outer compared to the middle and inner lagoons. The species composition reflects the open lagoon conditions in that several coastal marine species occurred in the lagoon. Some of the more abundant marine species (Table 3.5) included anchovies (Engraulidae), silversides (Atherinidae), spotted sand bass (Paralabrax maculatofasciatus), barred sand bass (P. nebulifei), queenfish (Seriphus politus), shiner surf perch (Cymatogaster aggregata), kelpfish (Heterostichus sp.), California halibut (Paralichthys califomicus), and diamond turbot (Hypsopsetta guttulata). Estuarine fish such as California killifish (Fundulus parvipinnis), gobies (Gobiidae), and pipefish also were collected. Mean total densities ranged from 0.016 to 7.90 fish per square meter (Table 3.5, Appendix B.2). Densities were lowest in the outer lagoon, intermediate in the middle lagoon, and highest in the east inner lagoon. Few fish were collected in the west inner lagoon during the summer survey, but moderate numbers were caught during the spring. Densities generally were higher in April than July for all lagoon sections, and were especially high in the east inner lagoon. Silversides (topsmelt and juvenile atherinids) and gobies were the most abundant fish caught during both surveys. The high densities in spring were due to recruitment of juvenile atherinids (probably topsmelt) and gobies. Silversides were primarily caught in the middle and east inner lagoon, with few collected in the west inner lagoon and none in the MEC Analytical Systems, Inc. 32 AGUA HEDIONDA LAGOON September 1995 Table 3.4.List of species collected by beam trawl, beach seine, and otter trawl at Agua Hedionda Lagoon for the July 1994 and April 1995 surveys. Species Mustelus califomicus Gymnura marmorata Engraulis mordax Anchoa compressa Engraulidae (< 50 mm) Fundulus parvipinnis Atherinops affinis Atherinidae (< 25 mm) Syngnathus sp. Syngnathus leptorhynchus Syngnathus aullscus Leptocottus armatus Paralabrax clathratus Paralabrax maculatofasdatus Paralabrax nebulifer Micropterus dolomieui Umbrina roncador Seriphus politus Cymatogaster aggregata Sphyraena argentea Hypsoblennius gentilis Heterostichus sp. Heterostichus rostratus Gobiidae Cleveland/a ios Acanthogobius flavimanus llypnus gilberti Quletula y-cauda Gillichthys mlrabllls Gobiidae (< 25 mm) Citharichthys stigmaeus Paralichthys califomicus Pleuronlchthys ritteri Hypsopsetta guttulata Svmohurus atricauda Common Name Grey smoothhound shark California butterfly ray Northern anchovy Deepbody anchovy Anchovy (< 50 mm) California killifish Topsmelt Atherinid (< 25 mm) Pipefish, unid. Bay pipefish Barred pipefish Staghom sculpin Kelp bass Spotted sand bass Barred sand bass Smallmouth bass Yellowfin croaker Queenfish Shiner surfperch California barracuda Bay blenny Kelpfish, juv Giant kelpfish Goby (unid.) Arrow goby Yellowfin goby Cheekspot goby Shadow goby Longjaw mudsucker Goby (< 25 mm) Speckled sanddab California halibut Spotted turbot Diamond turbot California tonquefish MEC Analytical Systems, Inc.33 v i t i f i f i r i r i r i r i r i i i • i • i iiii 1 Table 3.5.Mean total density per m2 of dominant taxa and total number of species of fish collected by beam trawl, beach seine, and otter trawl in different sections of Agua Hedionda Lagoon for the July 1994 and April 1995 surveys.0§ ! wI Species Engraulis mordax Anchoa compressa Fundulus parviplnnis Atherinops affinis Atherinidae (< 25 mm) Syngnathus sp. Syngnathus leptorhynchus Syngnathus auliscus Leptocottus armatus Paralabrax maculatofasciatus Seriphus politus Cymatogaster aggregata Hypsoblennius gentills Heterostichus rostratus Gobiidae Clevelandia ios Acanthogobius flavimanus llypnus gilbert! Quietula y-cauda Gillichthys mirabllis Qobiidae (< 25 mm) Parallchthys califomlcus Hypsopsetta guttulata Mean Total Density (m2) Total Number of Species Common Name Northern anchovy Deepbody anchovy California kllllflsh Topsmelt Atherinid (< 25 mm) Pipefish, unid. Bay pipefish Barred pipefish Staghom sculpin Spotted sand bass Queenfish Shiner surfperch Bay blenny Giant kelpfish Goby (unid.) Arrow goby Yellowfln goby Cheekspot goby Shadow goby Longjaw mudsucker Goby (< 25 mm) California halibut Diamond turbot July 1994 Outer Lagoon 0.0018 0.0013 0.0004 0.0007 0.0113 0.0013 0.0013 0.0013 0.0013 0.0275 11 Middle Lagoon 0.0060 0.0020 0.0180 0.9720 t 0.0068 0.0020 0.0040 0.0200 0.0016 0.0236 0.0040 0.0100 . 0.0040 , 0.0148 0.0220 1.1148 15 West Inner Lagoon 0.0060 0.0580 0.0013 0.0340 f 0.0020 0.0080 0.0102 0.0321 0.0028 L_ 0.0060 0.0580 . t , 0.0113 0.0022 0.2504 15 East Inner Lagoon 0.0320 0.0040 0.0600 1.2600 0.0020 0.0024 0.0020 0.5976 0.0040 0.0360 0.0980 0.0020 0.0080 0.0020 2.1220 16 April 1995 Outer Lagoon , . t , m . 0.0017 0.0117 0.0021 . 0.0159 4 Middle Lagoon 0.1220 0.7620 0.0920 0.0380 0.0697 0.0783 0.0180 0.0357 0.0520 0.0100 0.0260 0.0100 0.0400 0.1440 0.0527 0.1050 1.6580 18 West Inner Lagoon . 0.0620 0.0480 0.1580 0.0060 t 0.0080 0.0020 0.0018 t .. t 0.0380 0.0760 0.0020 0.0080 0.4292 0.0040 0.0440 0.8974 15 East Inner Lagoon 0.0020 0.0080 0.1400 3.2380 0.0060 0.0260 t 0.0140 0.1880 0.2200 t 0.0540 0.0420 3.8160 0.0040 0.1460 7.9040 14 1 1 ™ AGUA HEDIONDA LAGOON September 1995 m M outer lagoon. Topsmelt often is a dominant member of southern California lagoons and bays. They attach their eggs to aquatic vegetation such as eelgrass (Frey 1971), and high ** abundances commonly occur in lagoons when aquatic vegetation is prevalent. Larvae are " known to school in open shallow waters (Emmett et at. 1991), and this may relate to the high abundances of juvenile atherinids in the east inner lagoon. The highest abundance 5 of juvenile atherinids was collected at beach seine station S5 (Appendix B.2), located at the far end of the lagoon where the maximum depth is -2 ft MSL m Gobies were found in all sections of the lagoon during both surveys, although densities were highest for the east inner lagoon in April. Five species of goby were identified, the p most abundant being arrow goby and yellowfin goby. Arrow gobies (Clevelandia ios) were ^ found in all sections of the lagoon, but were most abundant in the middle and inner lagoon sections. They are most closely associated with muddy substrates (Brothers 1975). m H Yellowfin gobies (Acanthogobius flavimanus) were most abundant in the inner lagoon. This species is an introduced species that can displace native species, particularly other m gobies and sculpins (Usui 1981; Cross and Allen 1995). m After silversides and gobies, diamond turbot were the next most abundant fish. This p species was found in the middle and inner lagoon only, and in higher numbers in April m compared to July. Most of the fish collected in April were juveniles (Appendix B.4). """ California halibut occurred in low abundances in all sections of the lagoon, and had m approximately similar abundances in July and April (Table 3.5). More were caught in the middle compared to the outer and inner lagoons. «• 3.3.2 Sensitive Species "" The federally endangered tidewater goby (Eucyclogobius newberryi) once occurred in "" Agua Hedionda Lagoon. This species was not collected during the July 1994 or April 1995 ^ surveys. It is possible that the historical record may pre-date the construction of the lagoon "" (C. Swift, personal communication, 1994). The tidewater goby inhabits brackish waters, *" and would not be expected in the marine waters of the lagoon. 3.3.3 Historical Comparison _ Water quality characteristics of the lagoon were similar to that reported in previous studies (Bradshaw and Estberg 1973) in that temperatures were bit warmer, salinity was more ** variable, and visibility was lower in the inner lagoon compared to the outer and middle — lagoons. *"" Twenty-nine species of fish were collected during July 1994 and April 1995 surveys of p- Agua Hedionda Lagoon, which is generally lower than previous reports. Bradshaw and ^ Estberg (1973) listed a total of 22 species from one survey with beam trawls, interviews with fishermen, and observations. Bradshaw et al. (1976) reported a total of 42 species — from occasional surveys and from intake screen collections of the power plant. Recon la* M MEC Analytical Systems, Inc. 35 «• AGUA HEDIONDA LAGOON September 1995 urn m (1976) noted 44 species. SDG&E (1980) reported 54 species from monthly trawl, beach seine, and gill net collections. It is not known to what extent variations among reports may m relate to sampling effort and methodology versus changes in fish utilization of the lagoon •I over time. * In the present study, more fish were found further from the inlet as numbers were greater (i in the east inner lagoon. SDG&E (1980) also found that the highest abundances of fish occurred in the inner lagoon. IflJ ** Species composition in 1994 and 1995 was similar to that of previous studies in that the lagoon is utilized by coastal marine and estuarine fish. However, there are some notable 2 differences. Yellowfin goby, which were common in the inner lagoon during the 1994 and ™ 1995 surveys, have not been previously reported from the lagoon. L The abundances of California halibut in the present study were lower than in the past. In addition, the distribution of the species in the lagoon seems to have changed. California m halibut were mainly caught in the middle lagoon in the present study. California halibut m were one of the most abundant species reported by Bradshaw and Estberg (1973), and were only collected in the inner lagoon during their survey. Kramer (1990) documented PI that Agua Hedionda Lagoon was an important nursery ground for California halibut during ta 1987 and 1988, and reported highest densities in the inner and middle lagoons. -» Recent low abundances of California halibut in the lagoon have been confirmed by the n California Department of Fish and Game (CDFG), who has been conducting bimonthly surveys for halibut in four embayments in southern California, including Agua Hedionda '- Lagoon, since May 1994. They use a beam trawl for their surveys, and have rarely caught *» more than one halibut per trawl (C. Valley, CDFG, personal communication, 1995). "~ 3.4 Benthic Invertebrates Organisms living on or within aquatic sediments comprise the benthos. Invertebrates that "™ burrow or anchor within the sediment are termed infauna. These small animals, whose *• sizes range from microscopic to about 2 in were collected by hand cores in the subtidal _ stratum. Macroinvertebrates are larger and typically reside at or near the sediment surface; they were collected by beam trawl. Benthic infauna and macroinvertebrates are ta" a major food resource of bottom-feeding fish and diving birds. Invertebrates inhabiting _ shallows also are fed upon by wading birds and shorebirds. ""* A variety of benthic invertebrates live near, on, or in the subtidal sediments of Agua ,~ Hedionda Lagoon. A total of 143 taxa were collected by beam trawls and 76 taxa were collected by cores, with a combined total of 191 different taxa for both gear types from the *" July 1994 and April 1995 surveys. Results of the July 1994 and April 1995 — macroinvertebrate surveys, and the April 1995 infauna survey are presented separately below. MEC Analytical Systems, Inc. 36 m AGUA HEDIONDA LAGOON September 1995 *• Benthic data from the field surveys are given in Appendix C. Density and total counts of macroinvertebrates are presented in Appendix C.1 and Appendix C.2, respectively. ** Sediment data from the infauna cores are in Appendix C.3. Density data of benthic infauna ** are in Appendix C.4, and biomass of major taxonomic groups of infauna are given in Appendix C.5. ^M * 3.4.1 Species Composition, Density, and Distribution of Macroinvertebrates p A total of 143 invertebrate taxa were collected with beam trawls for both the July and April surveys. The number of species and density of macroinvertebrates were highly variable among individual trawls. There were 3 to 80 species collected per trawl, with total densities E ranging from 0.05 to 6.8 individuals per square meter (Appendix C.1). p Generally, more species were collected in the outer and middle lagoon sections, as H compared to the east inner lagoon during both the spring and summer surveys (Table 3.6). in July 1994, more species were collected in the west inner lagoon than in the east inner p lagoon. However, similar numbers of species were collected in April 1995 in the west and m east inner lagoon, and those numbers were lower overall than those collected in the middle and outer lagoons. m ta Abundances did not follow any clear pattern. Although densities of macroinvertebrates were fairly stable in the middle lagoon for the two surveys, large survey differences were "" seen for the other lagoon sections. Highest densities, due primarily to amphipods and tm bubble snails, were collected in the west inner lagoon during the July or summer survey. Mean densities were intermediate in the middle and outer lagoons, and were substantially p" lower in the east inner lagoon. In contrast, mean densities were highest in the east inner M lagoon during the April or spring survey, and lowest in the west inner and outer lagoons. Mean densities in the middle lagoon were fairly stable between surveys. — The most abundant macroinvertebrates included the cockle (Laevicardium substrlatum)', the mussel (Musculista senhousi); bubble snails (Acteocina inculta, Bulla gouldiana, """ Haminaea vesicula); mud dwelling snails (Alia carinata, Nassarius tegula); amphipod "• crustaceans (Amphithoe spp., Corophium spp., Ericthonius brasiliensis, Hyale sp., ^ Podocerus spp.); isopod crustaceans (Paracerceis sculpta); mysids; and shrimp (Crangon "~ nigromaculata, Hippolyte californiensis). w The species assemblages differed somewhat among lagoon sections. The cockle (L. substriatum) was more abundant in the outer and middle sections of the lagoon than in the "" inner lagoon. This is consistent with previous studies, and may relate to sediments being ... sandier in the outer and middle lagoon (Bradshaw and Estberg 1973). The mussel M. senhousi, which is an introduced species that may outcompete native clams in areas "* where it becomes established, was collected in relatively high densities in the east inner _ lagoon during both surveys. This species was not reported during previous surveys of the lagoon (Bradshaw and Estberg, 1973; Bradshaw et al. 1976). Crustaceans were "" seasonally abundant during the July 1994 summer survey throughout much of the lagoon — except for the east end, where they were collected in low abundances. Many of the ^ MEC Analytical Systems, Inc. 37 1 ) I I I 1 I I I I I I I I I 1 ii ii mm t i ii • i till I Table 3.6. Mean total density per m2 of dominant taxa, and total and mean number of species of macroinvertebrates collected by beam trawl in different sections of Agua Hedionda Lagoon for July 1994 and April 1995 surveys. Species Crepldula onyx (MO) Alia carinata (MO) Nassartus tegula (MO) Acteocina Inculta (MO) Su//a gouldiana (MO) Haminaea vesicula (MO) Musculista senhousl (MO) Laevicardium substriatum (MO) Te///na sp. B (SCAMIT) (MO) Tfteora /ubrfca (MO) Mysida (CFt) Paracarceis sculpta (CR) Ampithoe spp. (CR) Corophlum spp. (CR) Ericthonlus braslllensis (CR) Hya/esp. (CR) Podocerus spp. (CR) Periclimenes Infrasplnus (CR) Hippolyte califomlensls (CR) Crangon nlgromaculata (CR) Pagurus spp. (CR) Hamigrapsus oregonensls (CR) Dendraster excentricus (MP) TOTAL Number of species Mean number of species July 1994 Outer Lagoon 0.0680 t 0.0160 0.0040 t 0.7330 . 0.0020 . 0.0757 0.0060 .,0.0120 0.0080 0.0560 . .. 0.0673 1.5050 81 45 Middle Lagoon 0.0053 0.0213 0.0533 t 0.0433 0.0080 0.1460 t 0.0080 0,0767 0.0173 0.1920 1.0187 0.1013 0.0027 0.1533 t 0.0447 . 2.0360 38 14 West Inner Lagoon 0.0121 0.0750 t 0.6410 0.0090 0.0571 0.0033 0.4083 0.2000 0.1617 0.2467 0.9033 0.9200 0.0700 0.1557 0.0300 3.9869 30 14 East Inner Lagoon 0.0650 0.0240 0.0100 0.1850 0.0020 0.0040 0.0080 0.0050 t 0.0310 0.5300 0.0760 1.0290 19 7 April 1995 Outer Lagoon 0.0250 0.0500 0.0025 0.0325 0.0250 0.2050 0.0150 0.0275 0.0050 0.0025 . . 0.5250 39 24 Middle Lagoon 0.0272 0.0967 0.1794 0.0122 1.0928 0.0067 0.0022 0.3894 0.0256 t 0.0183 0.0689 0.0022 0.0956 2.2694 57 27 West Inner Lagoon 0.0025 0.0225 0.0050 0.0275 0.0150 0.1325 0.0450 0.0300 0.1325 0.0025 0.0100 0.5175 24 16 East Inner Lagoon 0.2700 0.0300 2.8100 0.2200 0.1900 0.1000 0.0050 0.0850 0.1100 0.0050 0.0250 4.0650 26 18 o § I !? Note: CR=crustacean, MO=mollusk, MP=minor phyla, PO=polychaete or oligochaete CD m m AGUA HEDIONDA LAGOON September 1995 abundant amphipod and isopod species have previously been reported to be common in the eelgrass beds of the lagoon (Bradshaw et al. 1976). The shrimp H. californiensis also is common in eelgrass beds (Morris et al. 1980). The lower abundances of these crustaceans in the east inner lagoon probably relates to the relatively poor development of eelgrass beds in that section of the lagoon. 3.4.2 Species Composition, Density, and Distribution of Infauna A total of 76 infaunal taxa were collected during the April survey. The number of species ranged from 2 to 25 per core, with total densities of 382 to 18,462 individuals per square meter. With the exception of stations with a high sand content (> 90% sand), there was little difference in assemblages from sediments ranging from 8 to 88% sand. Sediment data are presented in Appendix C.3, and count data by station are presented in Appendix C.4. Number of species, density, and biomass differed among vegetated and non- vegetated habitats and in areas of the lagoon where sedimentation has occurred. Generally, benthic populations were more diverse and abundant in the eelgrass beds than in non-vegetated sediments (Table 3.7). A notable exception was the populations in the eelgrass beds of the east inner lagoon, which are sparsely developed. There was little difference between the benthic community within or outside the eelgrass beds in that section of the lagoon. Other exceptions include areas where there has been deposition of littoral sands. The sand bars in the west inner lagoon are covered with eelgrass. On average, there were fewer species and individuals in the eelgrass beds on these bars than in most of the other eelgrass beds sampled in the outer, middle, and west inner lagoon sections. A final example concerns the eelgrass stations in the outer lagoon, which had substantially different benthic community characteristics (Appendix C.4). Station 1, in contrast to Station 2, had a poorly developed benthic infauna. Station 1 had one of the highest sand contents (93%) of any of the stations sampled. The high sand content at this station and the presence of the large sand bar on the west side of the lagoon provide evidence that there has been deposition of littoral sands at that station. Deposition of littoral sands is a constant problem for the outer lagoon, and maintenance dredging is conducted outside the eelgrass beds every one to two years. Benthic populations also were less developed in non-vegetated areas where there was littoral sand deposition. For example, there were few species and individuals of benthic infauna on the sand bar in the middle lagoon. The benthic infauna also was depauperate at Station 9 in the channel of the middle lagoon, where the sand content exceeded 90% (Appendices C.3 and C.4). Non-vegetated sediments with a lower sand content (8-88%) had moderate numbers of species and individuals in all lagoon sections. However, fewer species were collected on the sediment delta (bar) at the eastern end of the inner lagoon. Benthic assemblages at Stations 23 and 24 (Figure 2.3), which were located at depths of about -3 ft MSL, were more similar to those at deeper depths in the inner lagoon, than to Stations 25 and 26, which occurred at depths less than -2 ft MSL. Stations 25 and 26 had the lowest number of species and individuals of any of the sampled stations. MEC Analytical Systems, Inc. 39 1 1 i « I 1 I 1 I \ I 1 I I • I •1 11 • i mm o> i Table 3.7. Mean total density per m2 of dominant taxa and total number of species of benthic infauna collected by cores in different habitats and sections of Agua Hedionda Lagoon for the April 1995 survey. Species Nematoda (MP) Schlstomeringos rudolphl(PO) Leltoscoloplos pugettensls (PO) Polydora llgnl (PO) Polydora nuchalls (PO) Prlonospio haterobranchla (PO) Apoprionosplo pygmaea (PO) Splophanes mlsslonensls (PO) Pseudopolydora paucibranchiata (PO) Diplodmis sp. (PO) Capttella "capitata" (PO) Notomastus hemipodus (PO) Medlomastus sp. (PO) Mediomastus amblseta (PO) Oligochaeta (PO) Caecum callfomlcum (MO) Acteoclna Inculta (MO) Laevlcardium substriatum (MO) 7"e///na sp. (MO) Ertcthonlus brasiliensis (CR) Qrandldlerella laponlca (CR) PontoQenela rostrata (CR) Phoronida (MP) Mean Density (m2) Total Number of Species Mean Number of Species Mean Biomass (gm/m2) Outer Lagoon Eelgrass Bed 3374 64 318 764 64 955 318 •- - -• 446 191 828 637 64 m 10250 29 17 40 Non-Veg. t 127 64 . . 382 446 382 t 255 t 382 382 . t t 127 3310 16 10 30 Middle Lagoon Eelgrass Bed 2165 700 637 191 127 1719 6048 ( 127 191 318 64 13305 22 14 66 Non-Veg. t t 955 255 509 891 573 . 64 64 64 t f 64 4265 18 9 22 Bar 64 . - t t 891 127 t t 64 . t . 1273 6 4 1 West Inner Lagoon Eelgrass Bed 64 2101 p t 255 318 64 1401 64 573 5411 382 . 12032 23 15 73 Non-Veg. t 828 159 32 286 159 32 95 64 159 64 32 32 32 223 2833 24 10 19 Bar 1019 191 64 . . 637 127 4902 . 64 64 . 318 . 8340 18 11 48 East Inner Lagoon Eelgrass Bed 64 64 1464 . 64 64 127 . . 64 637 64 2737 11 6 4 Non-Veg. 191 127 509 127 64 . 509 . 1910 64 64 4138 16 10 20 Bar 191 382 637 64 668 2260 12 5 10 1 §§ Note: CR=crustacean, MO=mollusk, MP=minor phyla, PO=polychaete or oligochaete Ol AGUA HEDIONDA LAGOON _ September 1995 IP ** The most abundant infaunal invertebrates (Table 3.7) included nematodes, phoronids, oligochaetes, and polychaete worms (Capitella capitata, Mediomastusspp., Leitoscoloplos 1"; pugettensis, Polydora ligni, Spiophanes missionensis). Amphipod crustaceans (Ericthonius " brasiliensis, Grandidierellajaponica), cockles and clams (Laevicardium substriatum and Tellina sp.), and bubble snails (Acteocina inculta, Haminaea visicula) also were common in different areas of the lagoon. The species assemblages reflected differences in lagoon habitats. Nematodes and oligochaetes, which feed on organic matter (particularly vegetation), were most abundant in the eelgrass beds and on the vegetated bar of the west inner lagoon. Cockles and clams were most abundant in the outer lagoon where sediments are sandier. Amphipods were most abundant in the eelgrass beds of the middle and inner lagoon. Phoronids were most abundant on non-vegetated sediments in the inner lagoon. 3.4.3 Sensitive Species The speckled scallop (Argopecten circularis), which is a protected species, was collected from all three sections of the lagoon during 1972 (Bradshaw and Estberg 1973; reported as Aequipecten aequisculcatus). The CDFG studied this species in the lagoon from 1984 to 1986. Large numbers were collected in 1984, but they had virtually disappeared by 1986 (Haaker et al. 1986). Only 1 individual was collected in the middle lagoon during the July 1994 survey. 3.4.4 Historical Comparison The high number (1 91) of infauna and macroinvertebrate taxa (76 from cores) observed in the present study indicates that the benthic fauna is as or more diverse than it was in the 1970's. Bradshaw and Estberg (1973), sampled 36 stations in all three sections of the lagoon using a modified Hayward orange peel grab, and reported over 70 species. Bradshaw et al. (1 976) provided a more expanded list with over 1 50 species in the lagoon, with about 1 00 species from subtidal habitats. Despite the similarity in overall diversity between the present and historical studies, there are differences now in the benthic fauna that were not reported earlier. Bradshaw and Estberg (1973) documented that benthic species diversity, density, and biomass were similar throughout the lagoon. The only differences they noted concerned the species assemblages, which generally were dominated by clams in sandy sediments and by worms in sediments with a higher silt content. During the present study, these same patterns in species assemblages were noted, but more importantly, there were differences among lagoon sections and habitats, particularly in sediment depositional areas. In general, there has been a degradation of benthic habitat in areas that have received substantial deposition of littoral sands or terrigenous sediments. The sandbar in the middle lagoon supported a reduced fauna. The sandbars in the west inner lagoon, which are covered with eelgrass, had fewer species and numbers of individuals than other eelgrass beds in that section of the lagoon and in the middle and outer lagoon. MEC Analytical Systems, Inc. 41 »• AGUA HEDIONDA LAGOON _ September 1995 m m The east end of the inner lagoon has undergone substantial infilling from runoff since its original construction, and since the 1970's when the last benthic surveys were conducted. *" Results of the present study indicated that the benthic fauna of the eelgrass beds in the m east inner lagoon was less diverse and abundances were lower as compared to eelgrass beds in other lagoon sections. Furthermore, there were fewer species and individuals as P depths became shallower on the sediment delta at the east end.i m m:m m1 MEC Analytical Systems, Inc. 42 ** AGUA HEDIONDA LAGOON September 1995 m m 4.0 SUMMARY PI il Tidal action drives the circulation in the lagoon, and water quality generally was good throughout the lagoon. However, visibility was less, and temperature and salinity were p more variable in the inner lagoon where there has been deposition of fine sediments and II depths are shallower. There was evidence of reduced habitat quality in areas where substantial sedimentation has occurred. m H Ecological resources reflected the tidal conditions of the lagoon, in that marine species comprised a substantial proportion of the fish and benthic species assemblages, and m eelgrass was an important component of the habitat. Furthermore, water associated birds, If particularly shorebirds that forage on intertidal flats, were dominants at the lagoon. There were variations in these resources among the different lagoon sections and habitats. P • Eelgrass beds were best developed in the outer and middle lagoons. Eelgrass was limited to shallower depths in the inner lagoon, with the densest bed occurring on the sandbars H in the west inner lagoon. Eelgrass beds along the shoreline of the inner lagoon were less • dense and more patchy in distribution. Eelgrass beds comprised the most valuable habitat for benthic organisms in the lagoon. More species and higher densities generally occurred m in these vegetated sediments. Exceptions were in areas where there has been substantial • sediment deposition such as the sandbars in the west inner lagoon and near the sandbar in the outer lagoon. Sedimentation and associated turbidity in the east inner lagoon probably has contributed to the reduction in eelgrass and its present distribution being • restricted to shallow depths in that section of the lagoon. The sandbars in the outer lagoon, middle lagoon, and west inner lagoon, which become m exposed during low tides, were suboptimal habitats. Few benthic resources were «. associated with these areas, and they were of limited use as loafing areas for birds. The sediment delta that has formed over the years in the east inner lagoon ranged from poor ** to moderate benthic habitat depending on the degree of deposition. There were fewer •- benthic species and individuals as depths on the del^a became shallower. Bird use was very high in the east end of the inner lagoon. The east inner lagoon offers — and assortment of habitats ranging from intertidal flats, salt marsh, salt panne, and a, seasonal ponds, which in combination are found nowhere else in the lagoon. The intertidal flat at the east end was the most heavily used habitat of the entire lagoon, with high *"• abundances of foraging shorebirds. Gulls, coots, and ducks loafed on the sediment delta — and intertidal flats, when not foraging in the open water. The intertidal flat grades into the sediment delta, which becomes exposed at low tide. Shorebirds were commonly observed """ foraging on the sediment delta when it was exposed. Benthic infaunal species number «. and abundances were low on the sediment delta; benthic fauna were not sampled on the intertidal flats. Given the high shorebird abundances on the intertidal flats, it would be **" interesting to compare benthic populations on the flats to those on the sediment delta to *• determine relative value as foraging habitat to shorebirds. Shorebirds moved to salt marsh and salt panne habitats for loafing during high tides. The proximity of these loafing habitats MEC Analytical Systems, Inc. 43 " AGUA HEDIONDA LAGOON September 1995 m «* to foraging habitat probably contributes to the utilization of the flats by shorebirds at the east end of the lagoon. • Fish were most abundant in the east inner lagoon where shallow depths and warmer temperatures appeared to provide more optimal habitat for juvenile silversides and gobies. B Wading birds such as herons and egrets, which feed on fish and invertebrates, were more ™ commonly observed in the east inner lagoon. ig The lagoon provided limited to moderate habitat for special status species or species of special interest. California Brown Pelicans loafed on the aquaculture racks of the outer _ lagoon. A small population of Belding's Savannah Sparrow occurred in the salt marsh at II the east end of the lagoon. California Least Terns and Western Snowy Plover, which used to nest at the lagoon, were rarely observed. Only one live speckled scallop was collected • in the middle lagoon. California halibut were collected in low abundances, with most H collected from the middle lagoon. p Species diversity in the present study was higher or lower than previous reports depending HI on the resource. A total of 61 water associated bird species were observed in the present study as compared to 55 from past studies. A total of 29 taxa of fish were collected, which m was lower than most other reports of 42 to 54 species. A total of 191 taxa of benthic • invertebrates ranging in size from the small infauna to larger macroinvertebrates were collected, which was higher than the over 150 species reported previously. p M Ecological resources, in general, were similar to previous studies of the lagoon, although there were some notable exceptions. Most importantly there has been a reduction in the m eelgrass habitat. There also has been a reduction in the use of the lagoon by endangered *m species of birds such as California Least Tem and Western Snowy Plover. California halibut abundances were lower, particularly in the inner lagoon, as compared to previous "* studies. Benthic resources have been altered in areas where there has been substantial *• sediment deposition. Additionally, there are some introduced species that now occur in the inner lagoon that were not previously reported from the lagoon. These include *" yellowfin goby, and the mussel Musculista senhousi. The occurrence of introduced *•* species is of some concern because of their capacity to displace native species in areas „,. where they become established. MEC Analytical Systems, Inc. 44 AGUA HEDIONDA LAGOON September 1995 LITERATURE CITED Hi Backman, T.W. and D.C. Barilotti. 1976. Irradiance reduction: Effects on standing crops of the Eelgrass Zostera marina in a coastal lagoon. Marine Biology. 34:33-40.n I* Bradshaw, J.S. and G.N. Estberg. 1973. An ecological study of the subtidal marine life of Agua Hedionda Lagoon. Environmental Studies Laboratory of the University of San Diego. ffl Submitted San Diego Gas and Electric Co., Part 1, 99 p.; Part 2,123 p. Bradshaw, J.S. B. Browning, K. Smith, and J. Speth. 1976. The natural resources of Agua B Hedionda Lagoon. U.S. Fish and Wildlife Service. Coastal Wetland Series #16.109 pp. _ Brothers, E.B. 1975. The comparative ecology and behavior of three sympatric California j| gobies. University of California, San Diego, Ph.D. Dissertation, 368 pp. S Cross, J.N. and L.G. Allen. 1995. Fishes Chapter 9 in Ecology of the Southern California Bight: A Synthesis and Interpretation. M.D. Dailey J.J. Reish, and J.W. Anderson (eds.). University of California Press, Berkeley. 926 pp. m H Deile, N.W. 1994. Agua Hedionda Watershed dry weather storm water monitoring data. City of Carlsbad, Utilities and Maintenance Department. m Ellis, J.D. 1954. Final report: dredging Agua Hedionda Slough Encina Generating Station. San Diego Gas and Electric. jm M Emmett, R.L., S.L. Stone, S.A. Hinton, and M.E. Monaco. 1991. Distribution and Abundance of Fishes and Invertebrates in West Coast Estuaries, Volume II: Species Life pi History Summaries. ELMR Rep. No. 8. NOAA/NOS Strategic Environmental Assessments «•" Division, Rockville, MD> 329 pp. *"" Frey, 1971. California's living marine resources and their utilization, California Department **" of Fish and Game. *" Haaker, P.L., J.M. Duffy, K.C. Henderson, D.O. Parker. 1988. The speckled scallop, *" Argopecten circularis, in Agua Hedionda Lagoon, San Diego County, California. 32 pp. Jenkins, S.A. and D.W. Skelly. 1988. An evaluation of the coastal data base pertaining to """ seawater diversion at Encina Power Plant Carlsbad, CA. Submitted to San Diego Gas & ^ Electric. "* Jenkins, S.A. and J. Wasyl. 1993. Numerical modeling of tidal hydraulics and inlet closures *. at Agua Hedionda Lagoon. Submitted to San Diego Gas and Electric, Co. MEC Analytical Systems, Inc. 45 m AGUA HEDIONDA LAGOON September 1995 m H Kramer, S.H. 1990. Habitat specificity and ontogenetic movements of juvenile California Halibut, Paralichthys californicus, and other flatfishes in shallow waters of Southern p California. Southwest Fisheries Science Center, National Marine Fisheries Service, NOAA. Leeds, C.T. 1953. Memorandum regarding development of Agua Hedionda Lagoon. San p Diego Gas & Electric. B Leighton and Associates. 1988. Report of test results and laboratory analysis, San Diego Gas and Electric Encina Power Plant Dredge Material, Carlsbad, California. Project No. 4860085-02.m • Lillevang, O.J. 1966. Coastal power generation; its relationship to beaches. Shore and Beach: 34(2) :2-8. W^II ™ MEG Analytical Systems, Inc. 1993a. Field investigations for lagoon dredging and chemical analysis of sediments: data report. Submitted to San Diego Gas & Electric. ™m MEC Analytical Systems, Inc. 1993b. San Dieguito Lagoon restoration project regional _ coastal lagoon resources summary: Draft technical memorandum. Submitted to Southern M California Edison Company. m Morris, R.H., D.P. Abbott, and E.G. Haderlie (eds.). 1980. Intertidal Invertebrates of U California. Standford University Press, Standford, California. 690 pp. m Pike, P., SDG&E, personal communication, 1994. H Plumb, R.H. 1981. Procedures for Handling and Chemical Analysis of Sediment and •» Water Samples. Technical Report EPA/CE-81-1, prepared by Great Lakes Laboratory, ta State University College at Buffalo, Buffalo, N.Y., for the U.S. Environmental Protection Agency/Corps of Engineers Technical Committee on Criteria for Dredged and Fill Material. "" Published by the U.S. Army Engineer Waterways Experiment Station, CE, Vicksburg, Miss. lM Point Reyes Bird Observatory. 1992. Shorebird numbers in wetlands of the Pacific Flyway: "" A summary of counts from April 1988 to January 1992. ta> RECON. 1976. Environmental impact report No. 329, Agua Hedionda specific plan. *" Submitted to the City of Carlsbad. Ma Ritter, J.R. 1972. Cyclic sedimentation in Agua Hedionda Lagoon, Southern California. "* Journal of the Waterways, Harbors and Coastal Engineering Division, Proceedings of the *" American Society of Civil Engineers. 98(WW4): 597-602. "™ Swift, C. Los Angeles National History Museum, fish section, personal communication, 1994. MEC Analytical Systems, Inc. 46 M AGUA HEDIONDA LAGOON _ September 1995 iII Ultrasystems, Inc. 1983. Supplemental environmental studies. Submitted to City of Carlsbad. P H U.S. Army Corps of Engineers (USACOE). 1973. Flood plain information: Agua Hedionda Creek, Pacific Ocean to Buena San Diego County, California. Submitted to San Diego pj County. m U.S. Army Corps of Engineers (USACOE). 1994. Reconnaissance report: Pacific coast • shoreline, Carlsbad San Diego County, California.m Usui, C.A. 1981 . Behavioral, Metabolic, and Seasonal Size Comparison s of an Introduced B Gobiid Fish (Acanthogobius flavimanus) and a Native Cottid (Leptocottus armatus), from Upper Newport Bay, California. M.S. Thesis, California State University, Fullerton, CA. Valley, C. California Department of Fish and Game, personal communication, 1995. Wells, P.E. and D.W. Price. 1989. Agua Hedionda Lagoon, San Diego County: Sanitary m m survey 1985-1989. State Department of Health Services, Environmental Management Branch. MEC Analytical Systems, Inc. 47 •t'»'-r-j"«^fi '»* „ $*»*/•? v<t>;M-lt*t. „ ^,%-/'.-f'*,|k''L<->>"'•' 'i-Vt?1'-^ '-,* ."**>v' •• - .'' f'-^i^^^•^^g^^j^^iyilis^iii^i^^^i^^^^i m • A.1 Total Abundances by Species and Survey i i ii ii Species i i i r i r i r i i i i i i * a mm • i • i i i Agua Hedionda Lagoon Bird Data Total abundances by survey Aug94 . 30 1 4 1 1 13 1 5 Sep94 . 58 6 3 10 9 t t t 1 Oct94 18 7 3 63 12 3 6 2 46 7 19 Nov94 3 26 6 124 29 1 2 . . Dec 9 4 2 8 9 6 65 16 3 3 4 3 23 9 Jan95 5 19 20 9 9 29 2 3 1 . 77 175 53 2 Feb95 7 17 3 4 1 14 4 1 10 3 51 56 Mar 9 5 12 18 1 1 2 29 2 4 6 53 16 Apr 9 5 1 7 5 35 10 1 1 7 14 33 2 1 May 9 5 27 4 1 2 10 . 2 11 . Jun9( 52 6 2 1 20 21 8 12 1 1 300 10 25 25 21 80 11 Pacific Loon Horned Grebe Eared Grebe Western Grebe Clark's Grebe Pied-billed Grebe California Brown Pelican (FE,SE) Double-crested Cormorant (CSC) Great Blue Heron Green-backed Heron Great Egret Snowy Egret Brant Mallard Gadwall Northern Pintail Green-winged Teal Cinnamon Teal American Wigeon Northern Shoveler Scaup spp. Redhead Canvasback Lesser Scaup Buf flehead Surf Scoter Ruddy Duck Red-breasted Merganser Turkey Vulture Cooper's Hawk (CSC) Red- tailed Hawk Northern Harrier (CSC) Osprey (CSC) American Coot Semipalmated Plover Western Snowy Plover (FT, CSC) Killdeer Black-bellied Plover Long-billed Curlew (FC3,CSC) Whimbrel Spotted Sandpiper Greater Yellowlegs Willet Sandpiper, unidentified Least Sandpiper Dunlin Western Sandpiper Sanderling Dowitcher spp. Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened 55 1 83 134 147 1 1 1 59 121 2 70 1 8 94 4 93 12 3 26 8 2 1 104 , 19 104 3 1 2 58 363 84 5 t 51 4 101 . 27 98 4 1 1 43 601 65 49 22 1 62 110 6 18 45 1 2 2 38 99 45 296 . 6 1 1 1 119 23 40 2 24 16 483 1 125 80 1 52 2 71 550 1 2 1 1 111 62 53 4 66 2 15 145 8 1 83 25 46 1 3 68 63 22 5 342 13 4 1 1 46 10 4 12 1 5 44 116 28 165 1 3 30 51 1 4 4 1 14 4 576 1 . 3 1 2 1 1 2 5 1 2 2 i t i i i i i i i t i r i t i mm mm mm • i BI i ii Species Short-billed Dowitcher Marbled Godwit Common Snipe American Avocet Black-necked Stilt Red-necked Phalarope Western Gull California Gull (CSC) Ring-billed Gull Bonaparte's Gull Heermann's Gull Forster's Tern California Least Tern (FE,SE) Caspian Tern Black Skimmer (CSC) Mourning Dove Anna's Hummingbird Belted Kingfisher Black Phoebe Northern Rough-winged Swallow Barn Swallow Cliff Swallow Loggerhead Shrike (CSC) Marsh Wren Common Yellowthroat Western Meadowlark Red-winged Blackbird House Finch Savannah Sparrow Belding's Savannah Sparrow(FC2,SE) Lincoln's Sparrow Song Sparrow Common Raven American Crow Total Aug94 16 Agua Hedionda Lagoon Bird Data Total abundances by survey Sep94 Oct94 Nov94 Dec 9 4 Jan95 Feb95 Mar 9 5 Apr 9 5 May95 Jun95 38 6 42 2 1 7 3 12 g 2 14 4 . t 2 100 36 1 4 61 2 2 1 40 5 1 1 ...... 2 1 •^ — — — ^. 1260 20 33 46 5 10 6 1 1 1 1 1 2 1 12 3 10 8 1 1095 50 9 2 1250 693 2 1 1 1 2 1 5 2 2 3285 61 17 4 83 29 27 2 5 . 1 1 2 t . 7 s 4 . • 1391 37 75 16 10 2 2 6 10 13 2 6 32 t 4 1 1 3 6 1 2 . , . 8 8 1385 1262 34 16 2 . 11 11 14 1 3 6 1 5 9 12 11 15 11 1 945 27 38 5 43 35 4 1 1 29 2 13 5 . 4 2 6 13 1 . . 1083 17 38 9 10 6 18 5 5 5 14 1 16 210 5 23 9 98 1 1 5 5 2 2 12 4 5 10 6 2 5 4 323 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened m m myi^ | A.2 Counts by Lagoon Section and Habitat for Each Survey m |g Agua Hedionda Lagoon Bird Data Reconnaissance Survey _ East Inner Lagoon ^ Total Abundances by Survey Species JUN94 JUL94 H California Brown Pelican (FE, SE) 2 • White-faced Ibis (FC2, CSC) 2 Black-crowned Night Heron 1 B Great Blue Heron 1 Great Egret 1 Snowy Egret 115 6 Western Grebe 2 3EGadwall 3 Mallard 2 Black-bellied Plover 3 E Semi-palmated Plover 14 12 Killdeer 8 9 Whimbrel 2 6 Willet 10 32 B Greater Yellowlegs 2 Sandpiper, unidentified . 2560 Least Sandpiper 1 Western Sandpiper 3 H Dowitcher spp. . 30 • Marbled Godwit 8 American Avocet 20 20 m Black-necked Stilts 6 I California Gull (CSC) 6 I* Western Gull 1 Pointer's Tem 24 P* Bam Swallow 2 . Cliff Swallow 7 Black Phoebe 1 House Finch 2 '"* Lesser Goldfinch 4 ha Common Yellowthroat 2 Red-winged Blackbird 3 18 _ Belding's Savannah Sparrow (FC2, SE) 5 16 Song Sparrow 4 IM TOTAL 267 2712 IP» 1^ Special Status Codes: CSC = California Department of Rsh and Game, Species of Special Concern FC2 = Category 2 Candidate for Federal listing ^ FE Listed as Endangered by the Federal Government II SE = Listed as Endangered by the State of California Pi M Agua Hedionda Lagoon Bird Data Reconnaissance Survey East Inner Lagoon Counts by Lagoon Section and Habitat m HP | I 1i m IP Hi Habitat Shore Aerial Intertidal Rat East Intertidal Flat East Intertidal Rat East Intertidal Rat East Intertidal Rat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Rat East Intertidal Flat East Intertidal Rat East Intertidal Flat East Intertidal Rat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Rat East Intertidal Flat East Intertidal Flat East Intertidal Rat East Intertidal Rat East Intertidal Rat North Intertidal Flat North Intertidal Rat North Intertidal Rat North Intertidal Rat South Intertidal Flat South Intertidal Flat South Intertidal Rat South Intertidal Rat South Intertidal Flat South Intertidal Flat South Open Water South Open Water South Open Water South Open Water South Open Water South Salt Marsh North Salt Marsh North Salt Panne North Salt Panne North Salt Panne North Salt Marsh/Transitional North Salt Marsh/Transitional North Salt Marsh/Transitional North Salt Marsh/Transitional North Salt Marsh/Transitional South Salt Marsh/Transitional South Freshwater Marsh North TOTAL Species Snowy Egret Black-crowned Night Heron Great Blue Heron White-faced Ibis (FC2, CSC) Great Egret Snowy Egret Gadwall Mallard Semi-palmated Plover Killdeer Whimbrel Willet Greater Yellowlegs Least Sandpiper Western Sandpiper Sandpiper, unid. Dowitcher Marbled Godwit American Avocet Black-necked Stilts California Gull (CSC) Western Gull Forester's Tern Snowy Egret Black-bellied Plover Willet American Avocet Snowy Egret Black-bellied Plover Semi-palmated Plover Killdeer Willet Sandpiper, unid. California Gull (CSC) Western Grebe California Brown Pelican (FE, SE) Forester's Tern Bam Swallow Cliff Swallow Belding's Savannah Sparrow (FC2, SE) Song Sparrow Killdeer Whimbrel American Avocet Cliff Swallow House Finch Lesser Goldfinch Spong Sparrow Black Phoebe Common Yellowthroat Red-winged Blackbird JUN94 3 1 1 2 1 79 3 2 8 5 2 9 2 1 3 8 10+ 5 chicks 4 + 2 chicks 5 1 10 9 1 1 5 24 2 6 3 1 2 2 14 2 2 4 + 1 fledgling 2 5 2 4 2 1 2 3 267 JUL94 1 31 2325 30 12+ 5 chicks 1 5 1 235 16 9 5 3 18 2712 Special Status Codes: CSC = Callntertidal Flatornia Department of Rsh and Game, Species of Special Concern FC2 = Category 2 Candidate for Federal listing FE = Listed as Endangered by the Federal Government SE = Listed as Endangered by the State of California r ..... i r ........ i r ..... i r ..... i r ..... i i ....... i r i r~i mm mm mm m.m i i i i i Habitat Shore Aerial Aerial Aerial Aerial Aerial Aerial Acjuaculture racks West Bank West Bank West Bank West Bank West Beach North Beach South Beach South Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat August 1994 Species California Brown Pelican (FE,SE) Snowy Egret Mallard Mourning Dove Northern Rough-winged Swallow Barn Swallow California Brown Pelican (FE.SE) Great Blue Heron Western Gull California Gull (CSC) Ring-billed Gull Killdeer California Brown Pelican (FE,SE) Western Gull Great Blue Heron Snowy Egret Semipalmated Plover Killdeer Black-bellied Plover Greater Yellowlegs Willet Sandpiper, unidentified Least Sandpiper Dowitcher spp. Short-billed Dowitcher Marbled Godwit American Avocet Western Gull Forster's Tern Caspian Tern Great Blue Heron Green-backed Heron Great Egret Whimbrel Willet Marbled Godwit California Gull (CSC) Semipalmated Plover Killdeer Spotted Sandpiper Willet Least Sandpiper Western Sandpiper Marbled Godwit Inner Lagoon, west Outer Middle of Bristol Lagoon Lagoon Cove 3 5 . « 24 1 5 1 1 . * i Inner Lagoon, east of Bristol Cove . 5 12 5 2 2 3 31 1 89 3 52 2 43 351 74 46 16 32 6 3 5 3 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened i i i i [ i f i f i i i r i r i i i i i t i r i i i i i i i i i i i Habitat Shore Intertidal Flat West Intertidal Flat West Other perches North Other perches South Other perches South Open water Open water Open water Open water Open water Open water Riprap South Riprap South Salt Marsh North Salt panne North Salt panne North Salt panne North Salt panne North Salt panne North Salt panne North Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat August 1994 (continued) Species Brant Willet Western Gull Great Blue Heron Belted Kingfisher Double-crested Cormorant (CSC) Osprey (CSC) Forster's Tern Belted Kingfisher Northern Rough-winged Swallow Barn Swallow Willet Western Gull Belding's Savannah Sparrow(FC2,SE) Semipalmated Plover Killdeer Black-bellied Plover Sandpiper, unidentified Least Sandpiper Dowitcher spp. Total Outer Lagoon 1 1 Middle Lagoon Inner Lagoon, west of Bristol Cove Inner Lagoon, east of Bristol Cove 2 13 13 52 12 1 5 37 21 17 925 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened I 1 I I 1 I 1 I II ! I 1 I 1 I 1 I 1 f 1 I : I I I I • | [ 1 I I Habitat Shore Aerial Aquaculture racks Aquaculture racks Aquaculture racks Aquaculture racks Beach South Beach South Beach South Intertidal Flat East lAtertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat North Intertidal Flat North Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat West Other perches North Other perches North Other perches South Open water Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat September 1994 Species Great Egret California Brown Pelican (FE,SE) Double-crested Cormorant (CSC) Western Gull Forster's Tern Western Gull California Gull (CSC) Ring-billed Gull Great Egret Snowy Egret Green-winged Teal American Coot Semipalmated Plover Killdeer Black-bellied Plover Whimbrel Greater Yellowlegs Willet Sandpiper, unidentified Least Sandpiper Western Sandpiper Dowitcher spp. Marbled Godwit American Avocet Western Gull Forster's Tern Caspian Tern Whimbrel Willet Great Blue Heron Semipalmated Plover Whimbrel Spotted Sandpiper Willet Least Sandpiper Western Sandpiper Marbled Godwit House Finch Song Sparrow Willet Great Blue Heron Western Gull Belted Kingfisher Black-bellied Plover Outer Lagoon 58 6 8 3 1 2 . Inner Lagoon, west Middle of Bristol Lagoon Cove 1 1 3 1 1 2 1 1 1 1 1 2 Inner Lagoon, east of Bristol Cove 8 32 2 2 1 1 4 96 11 96 1 1 31 601 43 12 22 33 4 17 33 5 5 1 22 37 3 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened • i B i m i • i 11 11.11 Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat September 1994 (continued) Inner Inner Lagoon, west Lagoon, east Outer Middle of Bristol of Bristol Habitat Shore Species Lagoon Lagoon Cove Cove Open water Western Gull . . 1 2 Open water Heermann's Gull 1 . . Open water Forster's Tern 1 3 Riprap North Willet . . 1 1 Riprap South Willet . . 1 Salt Marsh North Belding's Savannah Sparrow(FC2,SE) ... 2 Salt panne North Great Blue Heron ... 1 Salt panne North Killdeer ... 16 Salt panne North Whimbrel ... 1 Salt panne North Willet ... 1 Salt panne North Common Snipe ... 1 Total 80 1 12 1161 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened i i i i i i i i i i i r i r i r i r i i i r i i i mm mm mmi iiii Habitat Shore Aerial Aerial Aerial Aerial Aerial Aerial Aerial Aerial Aerial Aquaculture racks Aquaculture racks Aquaculture racks Aquaculture racks Bank Bank Bank Bank Bank Bank Bank Bank Bank Bank Bank Bank Bank Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Other perches Other perches Other perches West North North North South South South South South South South South South South East East East East East East East East East South South South South South West Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat October 1994 Species Outer Lagoon Middle Lagoon Inner Lagoon, west of Bristol Cove Double-crested Cormorant (CSC) Great Egret 2 Northern Pintail Western Gull Forster's Tern Mourning Dove Black Phoebe House Finch Common Raven Double-crested Cormorant (CSC) 8 Western Gull 5 Forster's Tern 1 California Brown Pelican (FE,SE) 63 American Coot Spotted Sandpiper Willet Double-crested Cormorant (CSC) Great Blue Heron Snowy Egret Turkey Vulture Cooper's Hawk (CSC) Willet Least Sandpiper Western Gull House Finch Song Sparrow Great Blue Heron Semipalmated Plover Black-bellied Plover Least Sandpiper Dunlin Western Sandpiper Dowitcher spp. Marbled Godwit American Avocet Great Egret Snowy Egret Green-winged Teal American Wigeon Willet 1 Great Blue Heron 1 Mourning Dove House Finch American Crow Inner Lagoon, east of Bristol Cove Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened 26 3 1 1 1 2 16 30 2 1 57 45 83 45 296 4 20 33 2 1 19 2 r i r i r i r i r i r ri ri mm ii • i Habitat Shore Other perches Other perches Other perches Open water Open water Open water Open water Open water Open water Open water Open water Open water Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt panne Salt panne Salt panne Salt panne Salt panne Salt panne Salt panne Salt panne Salt panne North North South North North North North North North North North North North North North North North North Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat October 1994 (continued) Species Western Gull Belted Kingfisher Osprey (CSC) Eared Grebe Western Grebe Pied-billed Grebe Mallard Ruddy Duck American Coot Western Gull Ring-billed Gull Forster's Tern Long-billed Curlew (FC3,CSC) Willet Anna's Hummingbird Loggerhead Shrike (CSC) Marsh Wren Common Yellowthroat Great Egret Semipalmated Plover Western Snowy Plover (FT,CSC) Killdeer Greater Yellowlegs Dowitcher spp. Western Meadowlark Belding's Savannah Sparrow(FC2,SE) Song Sparrow Total Outer Middle Lagoon Lagoon Inner Lagoon, west of Bristol Cove Inner Lagoon, east of Bristol Cove 1 17 7 46 12 61 5 4 8 1 9 1 1 1 2 2 53 6 18 2 2 1 3 9 82 13 17 983 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened ri r i El B 1 • 1 1 I I i I Habitat Aerial Aerial Aerial Aerial Aerial Aerial Aerial Aerial Aerial Aquaculture racks Aquaculture racks Aquaculture racks Bank Bank Bank Bank Bank Bank Bank Bank Bank Bank Bank Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Other perches Open water Open water Open water Open water Open water Open water Open water Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat November 1994 Shore Species Double-crested Cormorant (CSC) Turkey Vulture Red-tailed Hawk Northern Harrier (CSC) Osprey (CSC) Whimbrel Willet Ring-billed Gull Mourning Dove California Brown Pelican (FE,SE) Double-crested Cormorant (CSC) Western Gull North Snowy Egret North Marbled Godwit North Western Gull South Pied-billed Grebe South American Coot South Black-bellied Plover South Willet South Dunlin South Marbled Godwit South Western Gull South Ring-billed Gull East Double-crested Cormorant (CSC) East Snowy Egret East Semipalmated Plover East Black-bellied Plover East Whimbrel East Willet East Dunlin East Western Sandpiper East Marbled Godwit East American Avocet East Western Gull East Ring-billed Gull East Forster's Tern North Belted Kingfisher Eared Grebe Western Grebe Pied-billed Grebe California Brown Pelican (FE,SE) Double-crested Cormorant (CSC) Scaup spp. Redhead Outer Middle Lagoon Lagoon 2 1 123 24 15 1 1 Inner Lagoon, west of Bristol Cove 10 2 4 1 9 1 Inner Lagoon, east of Bristol Cove 3 1 1 1 1 1 22 1 . . 45 1 1 1 34 20 2 1 23 39 1 15 483 14 40 360 1 1 25 4 1 1 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened 300 10 1 r i r i r i r r i r i r i r i t i ri K 1 M ft 1 I I I I Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat November 1994 (continued) Habitat Shore Species Open water Bufflehead Open water American Coot Open water Western Gull Open water Ring-billed Gull Open water Heermann's Gull Open water South Eared Grebe Open water South Ruddy Duck Open water South American Coot Sandbar Great Blue Heron Sandbar Willet Sandbar Marbled Godwit Sandbar Western Gull Sandbar Ring-billed Gull Salt Marsh North Marsh Wren Salt Marsh North Common Yellowthroat Salt Marsh North Western Meadowlark Salt Marsh North House Finch Salt Marsh North Savannah Sparrow Salt panne North American Avocet Salt panne North Black-necked Stilt Total Outer Lagoon 400 200 2 . . 300 60 Inner Lagoon, west Middle of Bristol Lagoon Cove 25 50 1 10 16 60 30 Inner Lagoon, east of Bristol Cove 400 2 25 24 . 2 1 5 2 2 1 2 1129 251 1904 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened 1 I 1 I 1 I I I 1 I ! I ! I 1 I 1 I I I 1 I I i 1 I 1 1 I I 1 | ] | 1 Habitat Shore Aerial Aquaculture racks Aquaculture racks Aquaculture racks Aquaculture racks Aquaculture racks Aquaculture racks Aquaculture racks Aquaculture racks Bank North Bank North Bank West Bank West Bank West Bank West Bank West Bank West Bank West Bank West Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat East Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat South Intertidal Flat South Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat December 1994 Species California Brown Pelican (FE,SE) California Brown Pelican (FE,SE) Double-crested Cormorant (CSC) Great Blue Heron Great Egret Snowy Egret Mallard Western Gull Heermann's Gull Sandpiper, unidentified Heermann's Gull Western Grebe Black-bellied Plover Whimbrel Willet Marbled Godwit Western Gull California Gull (CSC) Ring-billed Gull Double-crested Cormorant (CSC) Great Blue Heron Green-winged Teal Bufflehead American Coot Semipalmated Plover Killdeer Black-bellied Plover Sandpiper, unidentified Dowitcher spp. Marbled Godwit American Avocet Black-necked Stilt Forster's Tern Snowy Egret Semipalmated Plover Black-bellied Plover Whimbrel Willet Sanderling Marbled Godwit Western Gull Ring-billed Gull Black-bellied Plover Willet Inner Lagoon, west Outer Middle of Bristol Lagoon Lagoon Cove 3 58 5 1 2 2 3 42 1 80 1 6 1 1 2 8 4 6 2 Inner Lagoon, east of Bristol Cove 37 2 2 5 4 1 18 1 37 18 22 5 44 4 1 5 2 25 35 1 39 430 2 5 17 2 3 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened i i f i i i i i i i r I ! II II I ! I I II II II II II II III! Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat December 1994 (continued) Habitat Shore Species Intertidal Flat Other perches Other perches Other perches Other perches Other perches Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Riprap Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Seasonal Pond Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh South North North North South South South East East East East North North North North North North Forster's Tern Double-crested Cormorant (CSC) Western Gull California Gull (CSC) Forster's Tern Belted Kingfisher Horned Grebe Eared Grebe Western Grebe Pied-billed Grebe California Brown Pelican (FE,SE) Double-crested Cormorant (CSC) Gadwall Green-winged Teal Scaup spp. Bufflehead Ruddy Duck American Coot Western Gull California Gull (CSC) Snowy Egret Double-crested Cormorant (CSC) Great Blue Heron Great Egret Black-bellied Plover Willet Sandpiper, unidentified Marbled Godwit Ring-billed Gull Double-crested Cormorant (CSC) Gadwall Western Gull California Gull (CSC) Black-necked Stilt Northern Harrier (CSC) Anna's Hummingbird Marsh Wren Belding's Savannah Sparrow(FC2,SE) Song Sparrow Total Outer Lagoon 1 12 1 Inner Lagoon, west Middle of Bristol Lagoon Cove 1 2 1 1 2 3 6 1 3 2 21 21 44 5 100 1 7 1 1 1 1 5 40 6 3 . . . Inner Lagoon, east of Bristol Cove 11 4 1 6 , 2 12 17 8 2 1 1 2 7 4 162 84 496 649 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened i 1 I 1 I I I f I 1 I 1 I 1 I I I I I I I 1 I 1 I 1 I I I 1 I I I I I 1 I 1 Habitat Shore Aquaculture racks Aquaculture racks Aquaculture racks Aquaculture racks Bank Bank Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Other perches Other perches Other perches Other perches Other perches Other perches Other perches Other perches Open water Open water Open water Open water Open water Open water South West East East East East East East East East East East East East East East East East North North North North South South South South North North North North North North Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat January 1995 Species California Brown Pelican (FE,SE) Double-crested Cormorant (CSC) Willet Western Gull Willet Whimbrel Great Blue Heron Gadwall Green-winged Teal American Coot Semipalmated Plover Black-bellied Plover Willet Least Sandpiper Dunlin Western Sandpiper Dowitcher spp. Marbled Godwit American Avocet Western Gull California Gull (CSC) Ring-billed Gull Great Egret Snowy Egret Willet Marbled Godwit Great Egret Black-bellied Plover Willet Marbled Godwit Western Gull Forster's Tern Double-crested Cormorant (CSC) Great Blue Heron Osprey (CSC) Western Gull California Gull (CSC) Ring-billed Gull Horned Grebe Eared Grebe Western Grebe Pied-billed Grebe Double-crested Cormorant (CSC) Mallard Outer Middle Lagoon Lagoon 9 27 1 1 3 . 1 1 3 9 1 1 3 5 3 3 . Inner Lagoon, west of Bristol Cove 1 ( 1 4 . 1 2 1 3 1 1 2 1 1 11 14 3 1 2 Inner Lagoon, east of Bristol Cove 1 4 6 2 39 52 34 2 14 145 21 16 1 8 1 . 1 1 . 5 4 1 75 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened r i i i r i i i i i r i i i r i i i n i Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat January 1995 (continued) Habitat Shore Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Riprap Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Seasonal Pond Seasonal Pond Seasonal Pond Seasonal Pond Seasonal Pond Seasonal Pond Salt Marsh Salt Marsh Salt Marsh Salt panne North North North North North North North North North North North Species Gadwall Green-winged Teal Cinnamon Teal Redhead Canvasback Lesser Scaup Bufflehead Ruddy Duck Red-breasted Merganser Red-tailed Hawk American Coot Marbled Godwit California Gull (CSC) Ring-billed Gull Forster's Tern Belted Kingfisher Willet Semipalmated Plover Willet Dunlin Western Gull California Gull (CSC) Ring-billed Gull Gadwall Green-winged Teal Northern Shoveler Whimbrel Willet Black-necked Stilt Green-winged Teal American Coot Willet Gadwall Total Outer Lagoon Middle Lagoon I • I Inner Lagoon, west of Bristol Cove Bl 1 I i I I 2 17 18 23 33 1 8 1 1 23 1 1 1 3 1 58 46 25 36 6 1 1 Inner Lagoon, east of Bristol Cove 157 43 2 55 1 38 104 1 52 12 2 7 1 17 2 2 13 2 2 87 118 223 957 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened ii ii i i ii ii r i i i 11 ii r i i i • i B i • I ii 11 ii 11 Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat February 1995 Habitat Shore Species Aerial Aquaculture racks Aquaculture racks Aquaculture racks Aquaculture racks Bank Bank Bank Bank Bank Bank Bank Bank Bank Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat East North North North North North South West West East East East East East East East East East East East East East East East North North North North North North North North North North South South South Northern Rough-winged Swallow California Brown Pelican (FE,SE) Double-crested Cormorant (CSC) Great Blue Heron Snowy Egret Sandpiper, unidentified Western Grebe Double-crested Cormorant (CSC) Snowy Egret American Coot Spotted Sandpiper Spotted Sandpiper Ring-billed Gull American Crow American Coot Willet Clark's Grebe Great Blue Heron Snowy Egret Semipalmated Plover Black-bellied Plover Willet Least Sandpiper Dunlin Western Sandpiper Dowitcher spp. Marbled Godwit American Avocet Western Gull Ring-billed Gull Caspian Tern Snowy Egret Semipalmated Plover Whimbrel Willet Least Sandpiper Dunlin Western Sandpiper Sanderling Marbled Godwit California Gull (CSC) Western Grebe Clark's Grebe Snowy Egret Outer Lagoon Middle Lagoon Inner Lagoon, west of Bristol Cove Inner Lagoon, east of Bristol Cove 11 1 10 2 2 1 7 10 2 148 6 19 1 3 2 43 1 1 4 23 44 18 12 3 190 4 31 10 7 24 1 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened ii i i r i i • it ii r i Habitat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Other perches Other perches Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Riprap Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Seasonal Pond Seasonal Pond Seasonal Pond Seasonal Pond Seasonal Pond Salt Marsh Salt Marsh Salt Marsh Salt Marsh i r i r i r i i i i i Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat February 1995 (continued) • m § ii 11 ii Shore Species South Black-bellied Plover South Willet South Sandpiper, unidentified South Western Sandpiper South Marbled Godwit South Osprey (CSC) South Belted Kingfisher Eared Grebe Western Grebe Pied-billed Grebe Double-crested Cormorant (CSC) Great Blue Heron Gadwall Green-winged Teal Redhead Canvasback Lesser Scaup Bufflehead Surf Scoter Ruddy Duck Red-breasted Merganser American Coot West Sanderling Great Blue Heron Great Egret Snowy Egret Marbled Godwit Western Gull California Gull (CSC) Ring-billed Gull North Mallard North Gadwall North Green-winged Teal North Willet North Black-necked Stilt Willet North Marsh Wren North Common Yellowthroat North Belding's Savannah Sparrow(FC2,SE) Total Outer Lagoon t 4 1 . 1 2 2 1 7 . Inner Lagoon, west Middle of Bristol Lagoon Cove 1 1 22 61 4 23 1 1 2 1 5 1 1 1 1 . 7 51 18 65 2 28 . 1 1 1 2 3 1 6 . . Inner Lagoon, east of Bristol Cove 1 9 2 2 37 31 1 36 68 1 3 14 25 1 2 10 1 2 8 42 73 471 676 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened ti ti II ii it f i ii r i i i i i mm mm ft I El II II II Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat March 1995 Habitat Shore Aerial Aerial Aquaculture racks Aquaculture racks Aquaculture racks Bank North Bank North Bank North Bank North Bank North Bank North Bank North Bank North Bank North Bank North Bank North Bank North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat North Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Intertidal Flat South Species Northern Harrier (CSC) Common Raven California Brown Pelican (FE,SE) Double-crested Cormorant (CSC) Western Gull Mallard Gadwall Green-winged Teal Northern Shoveler American Coot Black-bellied Plover Whimbrel Least Sandpiper Western Sandpiper Dowitcher spp. Ring-billed Gull Caspian Tern Snowy Egret Gadwall Black-bellied Plover Long-billed Curlew (FC3,CSC) Whimbrel Willet Least Sandpiper Western Sandpiper Marbled Godwit American Avocet Western Gull Snowy Egret Gadwall Green-winged Teal American Coot Black-bellied Plover Whimbrel Willet Sandpiper, unidentified Least Sandpiper Western Sandpiper Sanderling Marbled Godwit American Avocet Western Gull California Gull (CSC) Ring-billed Gull Outer Lagoon 2 27 3 Middle Lagoon Inner Lagoon, west of Bristol Cove 13 Inner Lagoon, east of Bristol Cove 1 1 2 15 6 2 1 5 1 6 38 3 3 2 1 3 6 15 1 67 10 2 2 1. 26 3 1 1 1 3 116 10 54 1 9 2 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened II II II 1 ] f 1 f 1 I 1 I 1 I I II 1 II 1 II II Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat March 1995 (continued) Habitat Shore Species Other perches Other perches Other perches Other perches Other perches Other perches Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Riprap Seasonal Pond Seasonal Pond Seasonal Pond Seasonal Pond Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt panne Salt panne Salt panne Salt panne Salt panne North North North North West North North North North North North North North North North North North North North North North North North North Outer Middle Lagoon Lagoon Osprey (CSC) Forster's Tern Great Blue Heron Willet Western Gull Ring-billed Gull Eared Grebe Western Grebe Clark's Grebe Pied-billed Grebe Double-crested Cormorant (CSC) Mallard Gadwall Lesser Scaup Bufflehead Surf Scoter Ruddy Duck American Coot California Gull (CSC) Ring-billed Gull Caspian Tern Northern Rough-winged Swallow Barn Swallow Willet Snowy Egret Green-winged Teal Whimbrel ... American Avocet Snowy Egret Killdeer Willet Western Sandpiper Marsh Wren Common Yellowthroat Western Meadowlark Red-winged Blackbird Belding's Savannah Sparrow(FC2,SE) Song Sparrow Great Blue Heron Semipalmated Plover Killdeer Whimbrel Willet Inner Lagoon, west of Bristol Cove 1 1 1 1 2 3 10 14 1 1 9 7 74 17 43 6 6 1 . . Inner Lagoon, east of Bristol Cove 2 3 1 12 76 1 1 1 5 1 7 1 2 4 4 5 9 12 11 15 11 1 10 2 1 10 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened 1 3 1 I i 1 I I I a I I I 1 I I f 1 • I II KlBiKIftl ft ] 11 If I| Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat March 1995 (continued) Inner Inner Lagoon, west Lagoon, east Outer Middle of Bristol of Bristol Habitat Shore Species Lagoon Lagoon Cove Cove Salt panne North Least Sandpiper . . . 11 Salt panne North Western Sandpiper . . 2 Salt panne North American Avocet . . . 4 Salt panne North Black-necked Stilt . . . 2 Total 45 28 224 648 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened 11 ri • i ii r i r i * * mm mm t * mm mm mm mm m i i ii Habitat Shore Aerial Aerial Aguaculture racks Aquaculture racks Aquaculture racks Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Open water Open water East East East East East East East East East East East East East East East East East East East North North North North North North South South South South South South South South South South South West Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat April 1995 Species Willet Bonaparte's Gull California Brown Pelican (PE.SE) Double-crested Cormorant (CSC) Forster's Tern Double-crested Cormorant (CSC) Great Egret Snowy Egret Gadwall Semipalmated Plover Black-bellied Plover Whiinbrel Willet Dunlin Western Sandpiper Dowitcher spp. Marbled Godwit American Avocet Western Gull California Gull (CSC) Ring-billed Gull Forster's Tern Caspian Tern Black Skimmer (CSC) Pacific Loon Snowy Egret Whimbrel Spotted Sandpiper Willet Marbled Godwit Mallard American Coot Black-bellied Plover Whimbrel Willet Western Sandpiper Marbled Godwit Western Gull California Gull (CSC) Ring-billed Gull Forster's Tern Western Gull Eared Grebe Western Grebe Outer Lagoon 35 8 3 Middle Lagoon Inner Lagoon, west of Bristol Cove 1 1 1 2 3 2 2 50 6 6 10 1 2 Inner Lagoon, east of Bristol Cove 1 1 1 1 2 2 51 1 2 4 4 526 1 9 32 2 14 3 5 4 5 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened t i i i t i i i r i i i i i ii t i mm mm mm KIKJIIIIIIII Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat April 1995 (continued) Habitat Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Sandbar Seasonal Pond Seasonal Pond Seasonal Pond Seasonal Pond Seasonal Pond Seasonal Pond Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt panne Salt panne Salt panne Salt panne Salt panne Salt panne Salt panne Salt panne Shore Species Double-crested Cormorant (CSC) Mallard Gadwall Redhead Lesser Scaup Bufflehead Ruddy Duck Red-breasted Merganser American Coot Western Gull California Gull (CSC) Forster's Tern California Least Tern (FE,SE) Caspian Tern Great Blue Heron Snowy Egret Willet Marbled Godwit Western Gull California Gull (CSC) Heermann's Gull Forster's Tern Caspian Tern North Mallard North Gadwall North Green-winged Teal North Cinnamon Teal North American Avocet North Red-necked Phalarope North Marsh Wren North Common Yellowthroat North Red-winged Blackbird North Belding's Savannah Sparrow(FC2,SE) North Lincoln's Sparrow North Snowy Egret North Gadwall North Killdeer North Willet North American Avocet North California Gull (CSC) North Forster's Tern North Caspian Tern Total Outer Lagoon . 7 2 1 18 1 1 t ( Inner Inner Lagoon, west Lagoon, east Middle of Bristol of Bristol Lagoon Cove Cove 1 1 15 12 2 . 1 2 . 24 8 2 2 25 2 8 11 5 . • 1 1 1 3 4 3 1 6 . . 3 3 11 4 2 1 4 5 4 2 6 13 1 1 12 1 2 2 2 2 4 84 24 116 859 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened t i f i PI mm mm mm mm mm mm mm mm i mm mm Habitat Aquaculture racks Aquaculture racks Aquaculture racks Aquaculture racks Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Other perches Open water Open water Open water Open water Open water Open water Open water Open water Open water Open water Riprap Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt Marsh Salt panne Salt panne Salt panne Salt panne Salt panne Shore Species Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat May 1995 Outer Lagoon Middle Lagoon Inner Lagoon, west of Bristol Cove California Brown Pelican (FE,SE) 26 Double-crested Cormorant (CSC) 2 Red-breasted Merganser 1 Western Gull 31 East Double-crested Cormorant (CSC) East Great Blue Heron East Snowy Egret East Gadwall East Killdeer East Whimbrel East Willet East American Avocet East Western Gull East Black Skimmer (CSC) North Snowy Egret North Gadwall North Black-bellied Plover North Whimbrel South Spotted Sandpiper Forster's Tern California Brown Pelican (FE,SE) Double-crested Cormorant (CSC) Great Egret Snowy Egret 2 Western Gull Forster's Tern 2 California Least Tern (FE,SE) Cliff Swallow Common Raven American Crow North American Crow North Great Egret North Cliff Swallow North Marsh Wren North Common Yellowthroat North Red-winged Blackbird North Belding's Savannah Sparrow(FC2,SE) South Common Yellowthroat North Mallard North Gadwall North Killdeer North American Avocet North Red-winged Blackbird Total 64 1 1 1 3 2 2 15 Inner Lagoon, east of Bristol Cove 1 16 5 4 3 14 1 2 4 2 13 2 25 115 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened ii ii ii t i i 1 I I I 1 I ] I 1 I 1 1 I I I I ] I 1 ii i i ii i i Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat June 1995 Habitat Aquaculture racks Aquaculture racks Aquaculture racks Aquaculture racks Intertidal Plat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Intertidal Flat Other perches Open water Open water Open water Open water Open water Sandbar Sandbar Sandbar Sandbar Seasonal Pond Seasonal Pond Salt Marsh Shore Species California Brown Pelican (FE,SE) Double-crested Cormorant (CSC) Western Gull Heermann's Gull East California Brown Pelican (FE,SE) East Great Blue Heron East Snowy Egret East Mallard East Gadwall East Killdeer East Black-bellied Plover East Long-billed Curlew (FC3,CSC) East Whimbrel East American Avocet East Black-necked Stilt East Western Gull East California Gull (CSC) East Forster's Tern East Black Skimmer (CSC) North Marbled Godwit North Western Gull North American Crow South California Brown Pelican (FE,SE) South Green-backed Heron South Snowy Egret South Black-bellied Plover South Whimbrel South Willet South Marbled Godwit South Western Gull South Black Phoebe California Least Tern (FE,SE) Red-breasted Merganser Western Gull Forster's Tern California Least Tern (FE.SE) Cliff Swallow California Brown Pelican (FE,SE) Great Blue Heron Snowy Egret Western Gull North American Avocet North Black-necked Stilt North Mallard Inner Lagoon, west Outer Middle of Bristol Lagoon Lagoon Cove 23 6 10 1 . . . 4 1 4 1 1 2 1 1 1 1 1 4 2 1 2 2 1 . . 1 21 . 1 1 1 62 . 15 Inner Lagoon, east of Bristol Cove 6 1 17 19 7 2 4 1 1 13 3 5 1 2 2 . 1 1 3 2 6 2 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened I 1 I 1 1 I I 1 I 1 I 1 I 1 ! I f 1 I I I 1 I 3 I 1 I i I 1 I 1 I 1 I 1 I f Agua Hedionda Lagoon Bird Data Counts by lagoon section and habitat June 1995 (continued) Inner Inner Lagoon, west Lagoon, east Outer Middle of Bristol of Bristol Habitat Shore Species Lagoon Lagoon Cove Cove Salt Marsh North Gadwall ... 1 Salt Marsh North Cliff Swallow ... 9 Salt Marsh North Marsh Wren ... 4 Salt Marsh North Common Yellowthroat ... 5 Salt Marsh North Red-winged Blackbird ... 10 Salt Marsh North Belding's Savannah Sparrow(FC2,SE) ... 6 Salt Marsh North Song Sparrow ... 2 Salt Marsh North Common Raven ... 5 Salt panne North American Avocet ... 2 Total 126 18 28 151 Special status codes: CSC=California Department of Fish and Game, Species of Special Concern; FE=Listed as Endangered by the Federal Government; FT=Listed as Threatened by the Federal Government; SE=Listed as Endangered by the State of California FC2=Category 2 candidate for Federal listing for which USFWS does not have information to support listing FC3=USFWS category, too widespread or not threatened A.3 Total Abundances by Bird Category, Habitat, Lagoon Section, and Shore f i f i ri ri i i r i r i r i r i ri ri ii ii i i Location Outer Lagoon Outer Lagoon Outer Lagoon Outer Lagoon Outer Lagoon Outer Lagoon Outer Lagoon Outer Lagoon Habitat Open water Sandbar Intertidal Flat Bank Riprap Aquaculture racks Other perches Aerial Agua Hedionda Lagoon Bird Data Total Abundance August 1994 - June 1995 Bird Category Cormorants/Pelicans Skimmers/Terns Egrets/Herons Loons/Mergansers Grebes Coots/Dabbling Ducks/Geese Diving Ducks Gulls Cormorants/Pelicans Shorebirds Gulls Egrets/Herons Coots/Dabbling Ducks/Geese Shorebirds Gulls Egrets/Herons Grebes Shorebirds Gulls Passerines Egrets/Herons Shorebirds Gulls Cormorants/Pelicans Skimmers/Terns Egrets/Herons Loons/Mergansers Coots/Dabbling Ducks/Geese Shorebirds Gulls Skimmers/Terns Cormorants/Pelicans Egrets/Herons East Shore . 2 West North South Shore Shore Shore « . . . 1 . 1 1 3 . 1 8 1 1 6 15 21 2 Othe: 3 14 2 3 14 4 60 604 21 1 442 . . 87 457 7 7 1 3 1 117 r i i r i r i r i r i r i • i mm mm mm 1 K I II Location Middle Lagoon Middle Lagoon Middle Lagoon Middle Lagoon Middle Lagoon Middle Lagoon Middle Lagoon Habitat Open water Sandbar Intertidal Flat Bank Riprap Other perches Aerial Agua Hedionda Lagoon Bird Data Total Abundance August 1994 - June 1995 Bird Category Cormorants/Pelicans Kingfishers Skimmers/Terns Loons/Mergansers Grebes Coots/Dabbling Ducks/Geese Diving Ducks Gulls Passerines Cormorants/Pelicans Egrets/Herons Shorebirds Gulls Egrets/Herons Loons/Mergansers Coots/Dabbling Ducks/Geese Shorebirds Gulls Passerines Egrets/Herons Grebes Coots/Dabbling Ducks/Geese Shorebirds Shorebirds Passerines Kingfishers Egrets/Herons Gulls Passerines Doves Cormorants / Pel icans Egrets/Herons Gulls East Shore • f • . West North Shore Shore _ ( , . 5 1 , 99 1 4 1 1 1 1 1 1 1 1 1 t . . South Shore . 4 2 28 2 1 . 2 2 , f . Othe: 2 1 7 1 12 8 112 4 11 1 1 37 15 10 . . . . 3 5 1 1 1 \ ri r i r i r i r i r i r i i i r i r i i i K* •• Ei Bi Ki i i • i i i : Agua Hedionda Lagoon Bird Data Total Abundance August 1994 - June 1995 * East West North South j Location Habitat Bird Category Shore Shore Shore Shore Other j West Inner Lagoon Open water Cormorants/Pelicans . . . .10 Kingfishers .... 1 Ospreys .... 1 Skimmers/Terns .... 4 I Egrets/Herons .... 2 j Grebes . . . .114 Coots/Dabbling Ducks/Geese .... 270 Diving Ducks . . • • 441 Shorebirds .... 8 Gulls . . . .19 Passerines . . . .25 West Inner Lagoon Sandbar Cormorants/Pelicans .... 1 Skimmers/Terns .... 6 Egrets/Herons . . . .11 Shorebirds . . . . 75 Gulls . . .115 West Inner Lagoon Intertidal Flat Cormorants/Pelicans 1 Skimmers/Terns . . . 3 . Egrets/Herons ..61. Grebes . . . 5 . Coots/Dabbling Ducks/Geese . . . 5 . Shorebirds . . 278 215 Gulls 42 36 Passerines . . . 2 . West Inner Lagoon Bank Cormorants/Pelicans 2 Grebes . . . 2 . Coots/Dabbling Ducks/Geese 11 Shorebirds . . 92 15 Passerines 5 West Inner Lagoon Riprap Shorebirds ..12. West Inner Lagoon Other perches Cormorants/Pelicans . . 3 . . Kingfishers ..14. Ospreys ... 1 1 Skimmers/Terns .... 5 Egrets/Herons ..21. Shorebirds . . 1 . . Gulls 24 West Inner Lagoon Aerial Cormorants/Pelicans .... 3 Egrets/Herons .... 5 Gulls .... 1 Passerines .... 1 r i ri r i r i r i r i i i r i r i r i i K •1 R 1 • • 11 II II Location Habitat East Inner Lagoon Open water East Inner Lagoon Sandbar East Inner Lagoon Intertidal Flat East Inner Lagoon Bank East Inner Lagoon Beach East Inner Lagoon Riprap East Inner Lagoon Other perches East Inner Lagoon Salt Marsh Agua Hedionda Lagoon Bird Data Total Abundance August 1994 - June 1995 Bird Category Cormorants/Pelicans Skimmers/Terns Loons/Mergansers Grebes Coots/Dabbling Ducks/Geese Diving Ducks Gulls Passerines Hawks/Vultures Cormorants/Pelicans Coots/Dabbling Ducks/Geese Gulls Cormorants/Pelicans Skimmers/Terns Egrets/Herons Grebes Coots/Dabbling Ducks/Geese Diving Ducks Shorebirds Gulls Cormorants/Pelicans Skimmers/Terns Egrets/Herons Coots/Dabbling Ducks/Geese Shorebirds Gulls Passerines Hawks/Vultures Cormorants/Pelicans Shorebirds Gulls Shorebirds Ospreys Skimmers/Terns Gulls Passerines Egrets/Herons Coots/Dabbling Ducks/Geese Shorebirds Passerines Hummingbirds Hawks/Vultures East Shore t 2 12 25 14 75 56 1 80 2 4697 490 West North Shore Shore . t . , 2 , , 5 103 2 . 2 26 79 3 2 • 1 1 . . 2 18 22 216 2 1 South Shore 2 24 25 . . 4 51 . 295 3 1 45 21 84 2 2 3 67 1 • 1 Othe: 3 33 2 59 562 766 423 3 1 43 • 2 1 1 10 r i r i r i r i r i i i i i r i r i ii m M E i •i i i 11 ti ii Location Habitat East Inner Lagoon Salt panne East Inner Lagoon Seasonal Pond East Inner Lagoon Aerial Agua Hedionda Lagoon Bird Data Total Abundance August 1994 - June 1995 Bird Category Skimmers/Terns Egrets/Herons Coots/Dabbling Ducks/Geese Shorebirds Gulls Passerines Egrets/Herons Coots/Dabbling Ducks/Geese Shorebirds Ospreys Skimmers/Terns Egrets/Herons Coots/Dabbling Ducks/Geese Shorebirds Gulls Passerines Doves Hawks/Vultures East Shore West Shore North Shore 6 5 20 263 2 15 1 86 56 South Shore Other 1 1 8 12 3 23 25 14 6 B.1 Water Quality Data During Fish Surveys Agua Hedionda Lagoon Water Quality Data July 1994 Lagoon Sample Section Station Depth (m) Outer O1 B2 Middle B3 B4 B5 West Inner B6 B9 East Inner B7 B8 Temperature (deg. C) pH 0.5 2.1 4.3 0.5 1.6 3.3 0.3 0.7 1.4 0.3 0.9 1.9 0.3 1.1 2.2 0.3 1.2 2.4 0.3 1.1 2.1 0.3 0.9 1.5 0.3 0.8 1.7 21.30 20.94 20.83 20.89 21.07 21.08 23.14 23.14 23.14 23.77 23.80 22.87 22.71 22.45 22.34 23.00 23.11 22.43 23.19 22.15 22.11 24.72 24.81 24.73 24.74 24.75 24.68 7.71 7.73 7.73 7.74 7.74 7.74 7.75 7.76 7.76 7.73 7.77 7.86 7.80 7.82 7.85 7.74 7.74 7.77 7.71 7.75 7.75 7.67 7.69 7.69 7.68 7.70 7.70 32.7 32.7 32.7 32.7 32.7 32.7 32.5 32.5 32.7 32.6 32.7 32.7 32.6 32.7 32.6 32.6 32.8 32.5 32.6 32.7 32.3 32.5 32.7 32.6 32.6 32.5 32.5 7.3 7.5 7.5 7.5 7.5 7.4 9.5 9.3 9.6 10.4 9.6 12.7 9.9 10.1 10.6 10.0 9.1 9.9 10.2 9.8 9.8 9.8 9.1 9.5 9.0 9.2 9.4 Salinity Dissolved Secchi Oxygen Depth (m) 4.50 5.00 1.40 2.00 2.25 2.00 2.25 0.75 1.25 Agua Hedionda Lagoon Water Quality Data April 1995 Lagoon Sample Section Station Depth (m) Outer O1 B2 Middle B3 B4 B5 West Inner O2 B6 B9 East Inner B7 B8 Temperature (deg. C) pH 0.0 2.1 4.2 0.0 1.9 3.7 0.0 2.1 4.1 0.0 0.6 1.1 0.0 0.7 1.3 0.0 1.3 2.6 0.0 1.6 3.0 0.0 1.6 3.2 0.0 1.5 3.1 0.0 1.4 2.4 15.54 15.38 15.32 15.40 15.40 15.30 15.82 15.80 15.82 15.57 15.38 15.45 15.48 15.41 15.41 16.80 16.70 16.40 16.70 15.20 14.80 15.90 15.20 14.90 16.90 16.80 16.00 16.20 16.20 16.30 m t • . f f . . • 7.85 7.85 7.95 7.87 7.85 7.82 7.86 7.85 7.82 7.89 7.90 7.83 7.83 7.83 7.81 31.2 31.2 31.3 31.0 31.1 31.6 31.3 31.2 31.3 31.3 31.3 32.0 31.6 32.1 32.0 29.1 29.4 31.3 22.9 30.4 31.5 30.0 30.6 31.1 28.2 28.8 31.7 22.0 29.1 30.0 18.2 17.7 17.6 16.6 16.4 16.2 17.8 17.5 18.1 16.5 17.1 17.1 18.4 16.9 17.3 8.8 9.3 10.8 8.5 8.5 9.5 8.2 8.5 8.8 8.5 8.5 7.9 9.0 9.1 9.1 Salinity Dissolved Secchi Oxygen Depth (m) 1.25 1.25 0.75 1.50 1.25 1.10 1.00 1.00 1.25 1.25 m m m m B.2 Mean Density by Gear Type, Lagoon Section, Station, and Survey m m mm m Agua Hedionda Lagoon Fish Data Density per m2 by Gear Type and Station Outer Lagoon July 1994 Syngnathus leptorhynchus Syngnathus auliscus Paralabrax clathratus Paralabrax maculatofasciatus Hypsoblennius gentilis Heterostichus sp. Heterostichus rostratus Clevelandia ios Quietula y-cauda Gillichthys mirabilis Paralichthys califomicus Pleuronichthys rttteri TOTAL Number of species Beam Trawl B1 B2 0.004 0.008 0.004 0.004 0.004 0.004 0.004 0.032 7.000 0.003 0.003 0.017 Otter Trawl O1 0.002 0.004 0.001 0.002 0.017 0.023 3.000 0.027 5.000 Middle Lagoon July 1994 Engraulis mordax Anchoa compressa Fundulus parvipinnis Atherinops affinis Syngnathus leptorhynchus Syngnathus auliscus Leptocottus armatus Seriphus politus Sphyraena argentea Hypsoblennius gentilis Heterostichus rostratus Gobiidae Clevelandia ios Quietula y-cauda Paralichthys califomicus Hypsopsetta guttulata TOTAL Number of species B3 Beam Trawl B4 B5 Beach Seine S1 0.004 0.008 0.012 2.000 0.016 0.008 0.072 0.016 0.004 0.116 5.000 0.004 0.090 2.700 0.010 0.008 0.004 0.016 3.000 0.040 0.030 0.030 2.900 6.000 S2 0.030 0.010 2.160 0.010 0.020 0.100 0.020 0.030 0.010 0.020 0.040 0.080 2.530 12.000 m mm pm mm Agua Hedionda Lagoon Fish Data Density per m2 by Gear Type and Station West Inner Lagoon July 1994 Anchoa compressa Atherinops affinis Syngnathus leptorhynchus Syngnathus auliscus Paralabrax nebulifer Micropterus dolomieui Umbrina roncador Seriphus politus Cymatogaster aggregata Hypsoblennius gentilis Heterostichus rostratus Gobiidae Clevelandia ios Acanthogobius flavimanus Paralichthys californicus Hypsopsetta guttulata TOTAL Number of species Beam Trawl B6 B9 0.007 0.012 0.004 0.003 0.016 2.000 0.010 2.000 Otter Trawl O2 0.002 0.001 0.001 0.005 0.006 0.001 0.016 6.000 Beach Seine S3 0.030 0.090 0.060 0.060 0.020 0.010 0.040 0.050 0.080 0.030 0.030 0.010 0.510 12.000 S7 0.200 0.110 0.010 0.060 0.010 0.030 0.260 0.020 0.700 7.000 East Inner Lagoon July 1994 Mustelus californicus Gymnura marmorata Engraulis mordax Anchoa compressa Fundulus parvipinnis Atherinops affinis Syngnathus auliscus Paralabrax nebulifer Seriphus politus Cymatogaster aggregata Gobiidae Clevelandia ios Acanthogobius flavimanus Quietula y-cauda Gillichthys mirabilis Paralichthys californicus Hypsopsetta guttulata TOTAL Number of species Beam Trawl B7 B8 0.012 1.496 0.072 1.508 2.000 0.072 1.000 S4 0.010 0.010 Beach Seine S5 0.010 0.260 0.010 0.030 0.010 0.180 0.090 0.010 0.020 0.010 0.640 11.000 0.100 0.010 0.120 3.000 S6 0.010 0.150 0.020 0.200 6.030 1.420 0.020 0.400 0.020 8.270 8.000 m, m m m m Agua Hedionda Lagoon Fish Data Density per m2 by Gear Type and Station Outer Lagoon April 1995 i Quietula y-cauda Gobiidae (< 25 mm) Citharichthys stigmaeus Paralichthys californicus TOTAL Number of species Beam Trawl B1 B2 0.005 0.035 0.005 0.045 3.000 0.000 Otter Trawl O1 0.001 0.001 0.003 2.000 IFm m m Middle Lagoon April 1995 Atherinops affinis Atherinidae (< 25 mm) Syngnathus sp. Leptocottus armatus Paralabrax maculatofasciatus Cymatogaster aggregate Hypsoblennius gentilis Heterostichus rostratus Clevelandia ios Acanthogobius flavimanus llypnus gilbert! Quietula y-cauda Gillichthys mirabilis Gobiidae (< 25 mm) Paralichthys californicus Pleuronichthys ritteri Hypsopsetta guttulata Symphurus atricauda TOTAL Number of species B3 Beam Trawl B4 0.140 0.033 0.007 0.007 0.187 4.000 0.013 0.007 0.013 0.040 0.040 0.113 5.000 B5 0.025 0.015 0.015 0.030 0.005 0.090 5.000 Beach Seine S1 S2 0.120 1.840 0.300 0.120 0.240 0.350 0.090 0.090 0.100 0.010 0.050 0.200 0.220 0.110 0.350 4.190 15.000 0.490 1.970 0.160 0.070 0.070 0.020 0.060 0.160 0.040 0.080 0.050 0.290 0.080 0.170 3.710 14.000 Agua Hedionda Lagoon Fish Data Density per m2 by Gear Type and Station p m West Inner Lagoon April 1995 Engraulidae (< 50 mm) Fundulus parvipinnis Atherinops affinis Atherinidae (< 25 mm) Syngnathus sp. Leptocottus armatus Paralabrax maculatofasciatus Cymatogaster aggregata Clevelandia ios Acanthogobius flavimanus llypnus gilbert! Gillichthys mirabilis Gobiidae (< 25 mm) Paralichthys califomicus Hypsopsetta guttulata TOTAL Number of species Beam Trawl B6 B9 0.065 0.010 0.075 2.000 0.010 0.030 0.040 2.000 Otter Trawl O2 0.052 0.009 0.001 0.062 3.000 Beach Seine S3 0.010 0.160 0.050 0.030 0.030 0.010 0.230 0.040 0.030 0.190 0.780 10.000 S7 0.300 0.080 0.740 0.010 0.010 0.180 0.150 2.020 0.010 0.030 3.530 10.000 East Inner Lagoon April 1995 Anchoa compressa Fundulus parvipinnis Atherinops affinis Atherinidae (< 25 mm) Syngnathus sp. Leptocottus armatus Cymatogaster aggregata Clevelandia ios Acanthogobius flavimanus Quietula y-cauda Gillichthys mirabilis Gobiidae (< 25 mm) Paralichthys califomicus Hypsopsetta guttulata TOTAL Number of species Beam Trawl B7 B8 0.010 0.020 0.350 0.010 0.390 4.000 0.010 0.390 0.400 2.000 S4 0.170 2.380 13.000 Beach Seine S5 0.010 0.020 0.140 0.110 0.030 0.110 0.070 0.200 0.800 0.220 0.170 0.330 0.010 0.490 15.960 . 0.180 0.140 0.030 0.040 10.300 0.250 27.400 9.000 S6 0.010 0.070 0.110 0.020 0.550 0.160 7.710 0.010 0.310 8.950 9.000 I m I m 1 | B.3 Total Counts by Gear Type, Lagoon Section, Station, and Survey Agua Hedionda Lagoon Fish Data Total Counts by Gear Type and Station Outer Lagoon July 1994 Beam Trawl Otter Trawl B1 B2 O1 Syngnathus leptorhynchus . 1 2 Syngnathus auliscus 1 Paralabrax clathratus . 1 4 Paralabrax maculatofasciatus . . 1 Hypsoblennius gentilis . . 2 Heterostichus sp. 2 . . Heterostichus rostratus . 5 16 Clevelandia ios 1 Quietula y-cauda 1 Gillichthys mirabilis 1 Paralichthys californicus 1 Pleuronichthys ritteri 1 TOTAL 8 7 25 Number of species 735 Middle Lagoon July 1994 Beam Trawl Beach Seine B3 B4 B5 S1 S2 Engraulis mordax ... . 3 Anchoa compressa ... . 1 Fundulus parvipinnis ... 9 Atherinops affinis . . . 270 216 Syngnathus leptorhynchus 141 . 1 Syngnathus auliscus ... 1 Leptocottus armatus ... . 2 Seriphus politus . . . . 10 Sphyraena argentea . . . . 2 Hypsoblennius gentilis . 2 . Heterostichus rostratus 2 18 2 . 3 Gobiidae .41 Clevelandia ios . . . 4 1 Quietula y-cauda ... . 2 Paralichthys californicus . 1 . 3 4 Hypsopsetta guttulata ... 3 8 TOTAL 3 29 4 290 253 Number of species 2 5 3 6 12 Agua Hedionda Lagoon Fish Data Total Counts by Gear Type and Station West Inner Lagoon July 1994 Anchoa compressa Atherinops affinis Syngnathus leptorhynchus Syngnathus auliscus Paralabrax nebulifer Micropterus dolomieui Umbrina roncador Seriphus politus Cymatogaster aggregate Hypsoblennius gentilis Heterostichus rostratus Gobiidae Clevelandia ios Acanthogobius flavimanus Paralichthys californicus Hypsopsetta guttulata TOTAL Number of species Beam Trawl B6 B9 Otter Trawl O2 3 1 4 2 3 2 2 1 1 6 7 1 18 6 Beach Seine S3 S7 3 9 6 6 2 1 4 5 8 3 3 1 51 12 20 11 1 6 1 3 26 2 70 7 East Inner Lagoon July 1994 Mustelus californicus Gymnura marmorata Engraulis mordax Anchoa compressa Fundulus parvipinnis Atherinops affinis Syngnathus auliscus Paralabrax nebulifer Seriphus politus Cymatogaster aggregate Gobiidae Clevelandia ios Acanthogobius flavimanus Quietula y-cauda Gillichthys mirabilis Paralichthys californicus Hypsopsetta guttulata TOTAL Number of species Beam Trawl B7 B8 3 374 18 377 2 18 1 S4 26 1 3 1 18 9 1 2 1 64 11 Beach Seine S5 10 1 12 3 S6 1 15 2 20 603 142 2 40 2 827 8 mH mm ma Agua Hedionda Lagoon Fish Data Total Counts by Gear Type and Station Outer Lagoon April 1995 Quietula y-cauda Gobiidae (< 25 mm) Citharichthys stigmaeus Paralichthys califomicus TOTAL Number of species Beam Trawl B1 B2 1 7 9 3 Otter Trawl O1 1 1 2 2 Middle Lagoon April 1995 Atherinops affinis Atherinidae (< 25 mm) Syngnathus sp. Leptocottus armatus Paralabrax maculatofasciatus Cymatogaster aggregate Hypsoblennius gentilis Heterostichus rostratus Clevelandia ios Acanthogobius flavimanus llypnus gilberti Quietula y-cauda Gillichthys mirabilis Gobiidae (< 25 mm) Paralichthys califomicus Pleuronichthys ritteri Hypsopsetta guttulata Symphurus atricauda TOTAL Number of species B3 Beam Trawl B4 B5 21 5 1 1 28 4 2 1 6 6 17 5 5 3 Beach Seine 18 5 S1 12 184 30 12 24 35 9 9 10 1 5 20 22 11 35 419 15 S2 49 197 16 7 7 2 6 16 4 8 5 29 8 17 371 14 1 I Agua Hedionda Lagoon Fish Data Total Counts by Gear Type and Station West Inner Lagoon April 1995 Engraulidae (< 50 mm) Fundulus parvipinnis Atherinops affinis Atherinidae (< 25 mm) Syngnathus sp. Leptocottus armatus Paralabrax maculatofasciatus Cymatogaster aggregata Clevelandia ios Acanthogobius flavimanus llypnus gilbert! Gillichthys mirabilis Gobiidae (< 25 mm) Paralichthys californicus Hypsopsetta guttulata TOTAL Number of species Beam Trawl B6 B9 13 2 15 2 1 3 4 2 Otter Trawl O2 58 10 Beach Seine S3 S7 69 3 1 16 5 3 3 1 23 4 3 19 78 10 30 8 74 1 1 18 15 202 1 3 353 10 East Inner Lagoon April 1995 Anchoa compressa Fundulus parvipinnis Atherinops affinis Atherinidae (< 25 mm) Syngnathus sp. Leptocottus armatus Cymatogaster aggregata Clevelandia ios Acanthogobius flavimanus Quietula y-cauda Gillichthys mirabilis Gobiidae (< 25 mm) Paralichthys californicus Hypsopsetta guttulata TOTAL Number of species Beam Trawl B7 B8 35 1 39 4 39 40 2 S4 1 2 14 11 3 11 7 20 80 22 17 33 17 238 13 Beach Seine S5 1 49 1596 18 14 3 4 1030 25 2740 9 S6 1 7 11 55 16 771 1 31 895 9 B.4 Fish Size Data ft I KI Kill II II II II IIII 11 II II II II II II Agua Hedionda Lagoon Fish Size Data July 1994 Species: Mustelus calforrtcus Counts by Size Class (cm) Section Station Total Weight (gms) S6 150 Species: Qymnura marmorata Section Station Total Weight (gms) S4 139 S5 260 Species: Engrauas mordax Section Station Total Weight (gms) C S2 1.4 S4 0.2 I S6 661 Species: Anchoa compressa Section Station Total Weight (gms) C S2 4.3 1 S3 32 S6 6.5 Species: Fundukis parvipinnis Section Station Total Weight (gms) C SI 6.5 1 SS 9.4 1 S6 3.7 Species: Atherinops atMs Section Station Total Weight (gms) C S1 76.62 C S2 1683.3 S3 8.3S4 65.4 S5 0.1 S6 176.69 S7 37.5 Species: Syngnathus leptorhynchus Section Station Total Weight (gms) C 83 0.4 C B4 3.68 C B5 1.19 C S2 1.7 B9 0.94 O B2 1.41 O O1 3.7 Species: Syngnalhus auRseus Section Station Total Weight (gms) C SI 1.5 S3 7.5 1 S4 2 S7 12.8 O B1 0.4 Species: leptocottus armatus Section Station Total Weight (gms) C S2 45 Species: Paralibrax clathratus Section Staton Total Weight (gms) 0 B2 24 O O1 83 Species: Paralabrax rnacuatofasdatus Section Station Total Weight (gms) O 01 500 Species: Paralabnu nebuHer Section station Total Weight (gmt) O2 44 S3 93.1 S4 77 S7 38 Total Count 1 Total Count 1 1 Total Count 3 1 15 Total Count 1 3 2 Total Count 9 10 20 Total Count 270 216 9 26 1 803 20 Total Count 4 1 1 2 1 2 Total Count1 6 1 11 1 Total Count 2 Total Count 1 4 Total Count 1 Total Count 2 6 3 1 0-0.9 1-1.9 2-2.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 Counts by Size Class (em) 0-0.9 1-1.9 2-2.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 Counts by Size Class (em) 0-0.9 1-1.9 2-2.9 6 5 3 10 10 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 19 191 3 2 1 1 1 2 202 66 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 Counts by Size Class (em) 0-0.9 1-1.9 2-2.9 3-3.9 3-3.9 3-3.9 i 2 3-3.9 3-3.9 2 3-3.9 26 6 i 109 1 3-3.9 3-3.9 3-39 3-3.9 3-3.9 3-3,8 4-4.9 4-4.9 4-4.9 3 13 4-4.9 4-4.9 4-4.9 20 36 1 3 97 1 4-4.9 4-4.9 4-49 4-4.9 4-4.9 4-4.9 5-5.9 5-5.9 5-5.9 5-5.9 1 5-5.9 1 1 5-5.9 7 554 12 35 10 5-5.9 5-5.9 5-5.9 5-5.9 5-5.9 5-5.9 6-6.9 8-6.9 6-6.9 8-6.9 6-6.9 1 8-6.9 7 26 1 6 31 7 8-6.9 1 8-6.9 6-6.9 6-8.9 6-6.9 8-6.9 1 7-7.9 7-7.9 7-7.9 7-7.9 1 1 7-7.9 7-7.9 2 1 12 1 7-7.9 7-7.9 1 7-7.9 1 7-7.9 7-7.9 7-7.9 1 8-6.9 6-8.9 8-69 8-8.9 1 8-6.9 8-8.9 5 i 8 8-8.9 8-6.9 1 8-8.9 8-8.9 1 8-8.9 8-6.9 1 2 1 9-9.9 10-10.9 11-11.9 9-9.9 10-10.9 11-11.9 9-9.9 10-10.9 11-11.9 9-9.9 10-10.9 11-11.9 1 1 9-9.9 10-10.9 11-11.9 9-9.9 10-10.9 11-11.9 29 18 16 1 86 4 9-9.9 10-10.9 11-11.9 1 2 9-9.9 10-10.9 11-11.9 i 9-9.9 10-10.9 11-11.9 9-9.9 10-10.9 11-11.9 1 1 2 9-9.9 10-10.9 11-119 9-9.9 10-10.9 11-11.9 1 1 . 1 1 1 12-12.9 12-12.9 12-12.9 12-12.9 12-12.9 12-12.9 10 1 13-12.9 1 12-12.9 1 1 12-129 1 12-12.9 12-12.9 12-12.9 1 11-13.8 14-14.9 15-15.9 18-18.9 17-17.9 16-18.9 19-19.9 20-20.9 28-28.9 37375 1 13-13.9 14-14.9 15-15.9 16-18.9 17-17.9 18-16.9 19-19.9 20-20.9 28-26.9 37-37.9 1 • • 1 13-13.9 14-14.9 15-15.9 18-16.9 17-17.9 18-16.9 19-19.9 20-20.9 26-26.9 37-375 13-13.9 14-14.9 15-15.9 18-16.9 17-17.9 16-18.9 19-19.9 20-20.9 26-28.9 37-37.9 13-13.9 14-14.9 15-15.9 18-16.9 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-373 13-13.9 14-14.9 15-15.9 18-16.9 17-17.9 16-18.9 19-19.9 20-20.9 28-26.9 37-375 6 ... .... . . 13-13.9 14-14.9 15-15.9 16-16.9 17-17.9 18-16.9 19-19.9 20-20.9 28-26.9 37-375 '. . 1 1 .... . . 1 . . .... . . 1 ... . 1 13-13.9 14-14.9 15-15.9 16-16.9 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-375 1 . . .... . 13. .... . 1 ... . . 2132 1 . . 13-139 14-14.9 15-15.9 16-16.9 17-17.9 16-16.9 19-19.9 20-20.9 26-26.9 37-375 13-13.9 14-14.9 15-15.9 18-16.9 17-17.9 16-16.9 19-19.9 20-20.9 26-26.9 37J75 13-13.9 14-14.9 15-15.9 16-16.9 17-17.9 16-16.9 19-19.9 20-20.9 26-26.9 37J75 1 13-13.9 14-14.9 15-15.9 16-18.9 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-375 I 1 II II B I II I 1 II I 1I I i i Agua Hedionda Lagoon Fish Size Data July 1994 Spedes: Mfcroptena dolomieul Section Station Total Weight (gms) S3 0.2 Spedes: Umbrkw roncador Section Station Total Weight (gms) 02 44 Spedes: Seriphus poltus Section Station Total Weight (gms) C S2 4.5 B7 1.34 S3 0.5 Spedes: Cymatogaster aggregate Section Station Total Weight (gms) S3 22 S4 32 Spedes: Sphyraena argentea Section Station Total Weight (gms) C S2 0.8 Species: HypsoMennhis gentis Section Station Total Weight (gms) C B4 16.14 O2 16 S3 63.5 O O1 36 Species: Heterostichus sp. Section Station Total Weight (gms) O B1 0.2 Species: Heterosltehus rosfratus Section Station Total Weight (gms) C 83 4.1 C B4 46.25 C B5 2.07 C S2 5.3 B8 2.89 B9 0.66 O2 17.4 S3 44.1 S7 49.3 O B2 15.2 O O1 55.1 Spedes: Gobtdae Section Station Total Weight (gms) C B4 0.19 C B5 0.2 B8 0.08 B7 51.42 B8 0.8 56 5.71 S7 0.3 Species: Cleveland* tat Section station Total Weight (gms) C S1 0.4 C S2 0.1 S8 0.28 S7 0.3 0 B1 0.1 Species: Acanlhogoblus flavimanus Section Station Total Weight (gms) S3 32.2 S4 115 1 S7 199.9 Total Count 2 Total Count 1 Total Count 10 3 1 Total Count 4 1 Total Count 2 Total Count 21 5 2 Total Count 2 Total Count 218 2 3 3 1 8 8 6 5 16 Total Count 4 1 1 374 18 142 1 Total Count 4 1 2 3 1 Total Count 3 18 26 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 2 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 5 5 1 2 1 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 2 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 4 1 1 173 159 42 17 1 138 4 1 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 4 1 1 . 1 3 1 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 1 4-4.9 4-4.9 4-4.9 4-4.9 4-4.9 2 4-4.9 4-4.9 4-4.9 1 i11 4-4.9 4-4.9 4-4.9 5-5.9 5-5.9 5-5.9 5-5.9 1 5-5.9 5-5.9 5-5.9 5-5.91 5 1 1 1 1 4 5-5.9 5-5.9 5-5.9 5 1 6-6.9 7-7.9 8-8.9 8-6.9 7-7.9 6-8.9 6-6.9 7-7.9 6-8.9 8-6.9 7-7.9 8-6.9 3 8-6.9 7-7.9 6-8.9 8-6.9 7-7.9 8-8.9 2 3 8-6.9 7-7.9 8-8.9 8-6.9 7-7.9 8-8.9 1 354 1 1 1 1 1 1 1 1 . 2 2 5 1 5 3 8-9.9 7-7.9 8-8.9 6-6.9 7-7.9 8-6.9 9-9.9 7-7.9 6-8.9 1 334 646 9-9.9 10-10.9 11-11.9 12-12.9 9-9.9 10-10.9 11-11.9 12-12.9 9-9.9 10-10.9 11-11.9 12-12.9 9-9.9 10-10.9 11-11.9 12-12.9 1 9-9.9 10-10.9 11-11.9 12-12.9 9-9.9 10-10.9 11-11.9 12-12.9 1 . . . 11 . 11 . 9-9.9 10-10.9 11-11.9 12-12.9 9-9.9 10-10.9 11-11.9 12-12.9 1 1 . . 1 2 1 2 1 1 2 '. . \ 9-9.9 10-10.9 11-11.9 12-12.9 9-9.9 10-10.9 11-11.9 12-12.9 9-9.9 10-10.9 11-11.9 12-12.9 1 . 1 . 1 . 1 . 15.1 13-13.9 14-14.9 15-15.9 19-18.9 17-17.9 16-18.9 19-19.9 20-20.9 29-25.9 37-374 13-13.9 14-14.9 15-15.9 16-15.9 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-37.9 1 ... ... . . 13-13.9 14-14.9 15-15.9 16-16.9 17-17.9 16-18.9 19-19.9 20-20.9 26-28.9 37475 13-13.9 14-14.9 15-15.9 18-18.9 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-375 13-13.9 14-14.9 15-15.9 16-16.9 17-17.9 18-18.9 19-19.9 20-20.9 29-26.9 37-379 13-13.9 14-14.9 15-15.9 19-19.9 17-17.9 18-18.9 19-19.9 20-20.9 28-28.9 37-37-9 13-13.9 14-14.9 15-15.9 16-16.9 17-17.9 16-18.9 19-19.9 20-20.9 26-26.9 37-375 13-13.9 14-14.9 15-15.9 16-16.9 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 3737.9 13-13.9 14-14.9 15-15.9 19-18.9 17-17.9 16-18.9 19-19.9 20-20.9 28-26.9 37-375 13-13.9 14-14.9 15-15.9 16-16.9 17-17.9 16-18.9 19-19.9 20-20.9 28-28.9 37-375 13-13.9 14-14.9 15-15.9 16-16.9 17-17.9 18-18.9 19-19.9 20-20.9 28-26.9 37-375 1 I I 1 I 1 I 1 I 1 1 1 I i I 1 I 1 I 1 I ] I i I 1 II f I II 11 I I II Agua Hedionda Lagoon Fish Size Data July 1994 Species: Quletula y-cauda Section StaBon C S2 S4 S6 O B1 Total Weight (gms) 0.71 2.28 0.1 Total Count 29 40 1 Species: QHchlnys mirabWs Section Station S4 0 Bt Total Weight (gms) 10.1 Total Count 11 Species: Parallchthys ealfomlous Section StationC B4 C SI C S2 02 S3 S4 S6S7 0 B1 Total Weight (gms) 1.84 53 14.9 124.7 19.3 75 125 150.2 Species: Pleuronlchtnys ritteriSection Station Total Weight (gms) 0 B1 41 Species: Hypsopsetta guttulata Section Staton C SI C S2 O2S3 S4 Total Weight (gms) 32 228.1 19165 7 Total Count 1 3 4 7 3 2 22 1 Total Count1 Total Count 3 9 11 1 Counts by Size Class (cm) 0-0.9 1-1.9 5 39 1 2-2.9 3-3.9 4-4.9 5-5.9 6-6.9 1 1 ... 4 .... 1 .... 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 13-13.9 Counts by Size Class (cm) 0-0.9 1-1.9 1 2-2.9 3-3.9 4-4.9 5-5.9 6-6.9 1 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 13.13.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 4-4.9 5-5.9 6-6.9 1 1 1 2 1 1 .... Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 4-4.9 5-5.9 6-6.9 7-7.9 6-8.9 9-9.9 10-10.9 11-11.9 12-12.9 ! . ii ... 12. 31 1.1. 11 11.. 7-7.9 6-8.9 9-9.9 10-10.9 11-11.9 12-12.91 13-13.9 1 1 13-13.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 4-4.9 5-58 6-6.9 '. '. '. 1 3 '. . . 1 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 3 ... 1.1. . 1 1 . . . 13-13.9 14-14.9 15-15.9 14-14.9 15-15.9 14-149 15-15.9 17-17.9 16-16.9 19-19.9 20-20.9 16-16.9 19-19.9 20-20.9 17-17.9 18-18.9 19-19.9 20-20.9 29-26.9 37-375 26-26.9 37-375 26-26.9 37-37.9 14-14.9 15-15.9 14-14.9 15-15.9 16-16.9 1 17-17.9 18-18.9 19-19.9 20-20.9 19-19.9 20-20.9 26-26.9 37-373 26-26.9 37-37:9 IlliVllilliilillllflllilKl I 1 II II Agua Hedionda Lagoon Fish Size Data April 1995 Spedes : Anchoa compressa Section Station Total Weight (gms)S4 3.5 Species: Engrauldae (< 50 mm) Section Station Total Weight (gnu) 1 O2 U Species: Fundukis pirvferrts Section Station Total Weight (gmi) SJ 7 S4 3.2 S5 0.3 SS 4 S7 155 Species: Alherinops aflW* Section Station Total Weight (gnu) C S1 0.85 C S2 35.95 S3 3.6 S4 3 S5 12.6S9 1.4 S7 2 Species: Alherinldae (< 25 mm) Section Station Total Weight (gms) C S1 13.45 C S2 23.5B7 0.05 S3 0.7 S4 0.7S5 52.9 se 02 S7 3.1 Species: Syngnalhus sp. Section Station Total WeigM (gms) C S1 14 C S2 5.65 S3 2.1 I S4 1.«5 Species: Leptocottus armatus Section Station Total Weight (gms) C S1 85.2 C S2 57.6S3 21 1 S4 48.5 1 S6 1.6 I S7 4.5 Species: Paralabrax macuUtofasclatus Section SUtlon Total Weight (gms)C B4 93.3 C BS 18 C S1 89.4 C S2 23.7 S7 0.1 Species: Cymatogaster aggregata Section Station Total Weight (gms) C B4 2.5 C BS 5 C S1 86.7 C S2 0.65 O2 49 S4 10.1 Species: HypsobtennKis genius Section Station Total Weight (gms) C St 10.3 Total Count1 Total Count 58 Total Count 1 21 1 30 Total Count 12 49 18 1449 7 8 Total Count 184 1971 5 11159S 11 74 Total Count 30 16 3 3 Total Count 12 7 311 21 Total Count2 5 24 7 t Total Count 1 3 35 2 10 7 Total Count 9 Counts by Size Class (em) 0-0.9 1-1.9 2-2.9 Counts by Size Class (cm)0-0.9 1-1.9 2-2.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 1 Counts by Size Class (cm)0-0.9 1-1.9 2-2.9 3 It 14 1230 4 2 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 1 Counts by Size Class (cm)0-0.9 1-1.9 2-2.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 '. t Counts by Size Class (em) 0-0.9 1-1.9 2-2.9 4 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 3-3.9 1-3.9 1 3-3.9 1 6 2 2 193 5 3-3.9 3-3.g 3-3.9 3-3.9 1 1 1 3-3.9 1 29 2 7 5 3-39 3 4-4.9 4-4.9 4-4.9 1 1 4-4.9 22 4-4.9 4-4.9 2 - 1 4-4.9 15 2 4-4.9 i 6 1 4-4.9 1 1 2 2 4-4.96 5-5.9 5-5.9 5-5.9 5 5-5.9 10 5-5.9 5-5.9 1 5-5.91 2 5-5.9 10 3 5-5.9 t 5-5.9 6-6.9 6-6.9 6-6.9 1 i 19 6-6.9 6-6.9 6-6.9 34 1 6-6.95 21 2 1 9-6.91 2 5 1 6-6.9 6-6.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 1 ... 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 5 ... 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 2342 1 2 3221 2 . 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 33.. 221. 1 . . . 11. . . 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 1 ... 2 ... . . 1 ... . . 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 2 . . . 1 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 13-13.9 14-14.9 15-15.9 16-16.9 13-13.9 14-14.9 15-15.9 16-16.9 13-13.9 14-14.9 15-15.9 16-16.9 13-13.9 14-14.9 15-15.9 18-16.9 13-13.9 14-14.9 15-15.9 16-16.9 13-139 14-14.9 15-15.9 16-16.9 13.1 1 1 2 13-13.9 14-14.9 15-15.9 16-16.9 13-13.9 14-14.9 15-15.9 16-16.9 13-13.9 14-14.9 15-15.9 16-16.9 13-13.9 14-14.9 15-15.9 16-18.9 17-17.9 18-18.9 19-19.9 20-209 26-26.9 37-37.9 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-37.9 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-37.9 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-372 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-37.9 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-373 3 . . 1 . . 1.1. . . 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-37.9 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37-372 17-17.9 18-18.9 19-19.9 20-20.9 26-26.9 37472 17-17.9 18-18.9 19-19.9 20-20.9 28-26.9 37-372 I 1 I I I 1 I 1 I } I 1 I ] I I I 1 I I I 1 I 1 I 1 I 1 I 1 I J I i I I Agua Hedionda Lagoon Fish Size Data April 1995 Species: Heterostichus rostratus Section Station Total Weight (gms) C B4 0.35C B5 0.15 S1 S2 3.3 1.7 Species: Cleveland los Section Station Total Weight (gins)S1 S2 B8 S3 S4 S5 S8 S7 3 4.2 0.2 0.2 4.9 4 8.4 3.4 Species: Acanftogobhis flavimanus Section Station Total Weight (gms) S1 S2 S3 S4 S5 S9 S7 1.5 2.9 35.1 73.9 3.9 16.1 8.8 Species: llypnus gflberti Section Station Total Weight (gms) C St 1,7 C S2 2.8 I B9 0.2 Species: Quietula y-eauda Section Station Total Weight (gms) C S2 1.3 I B7 0.5 I S4 7.3 I S5 1.1 O B1 0.3 Species: GJtichthyl mirabls Section Station Total Weight (gms) S1 S3 S4 S5 10.1 2 15.1 1 Species: OobMae (< 25 mm)Section Station Total Weight (gms) C C C C C D B3 B4B5S1 S2 B9 B7 B8B9 02 S3 S4 S5 SB S7B1 1.4 0.30.4 2.65 2.50.5 2 2.10.2 0.05 0.2 3.75 52.6 72.39 0.5 Species: Cltharichthys stlgmaeus Section Station Total Weight (gms) O O1 7 Total Count 23 9« Total Count10 191 1 20 18 55 IB Total Count1 4 23eo 141«IS Total Count5 81 Total Count5 2 22 31 Total Count 20 4 17 4 Total Count 216 6 22 2913 3539 31 3 33 1030 771 2027 Total Count1 Count! by Size Class (cm) 0-0.9 1-1.9 2-2.9 1 Counts by Sizt Class (em) 0-0.9 1-1.9 2-2.99 3 t 117 14 50 17 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 '. . t 1 5 8 3 Counts by Slz« Class (cm) 0-0.9 1-1.9 2-2.93 3 1 Counts by Size Class (em) 0-0.9 1-1.9 2-2.9 32 11 ! '. i Counts by Size Class (cm) 0-0.9 1-18 2-2.9 7 t 3 t 2 Counts by Size Class (cm)0-0.9 1-1.9 2-2.9 ! 2 '. t Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 4-4.9 5-S.9 6-9.9 7-7.9 9-B.9 9-9.9 10-10.9 11-11.9 12-12.9 1.1..... 3 ...... 5 3 1 ..... 4.1..... 3-3.9 4-4.9 5-5.9 <-t.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 1 ..... 13 ....... 3 '. '. '. '. '. '. . '. 4 ....... 5 ....... 1 ....... 3-3.9 4-4.9 5-5.9 9-6.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 1 ...... 2.1..... 3 10 8 1 .... 37 26 10 2 .... 9 ....... 0 9 2 2 ...8 4 ...... 3-3.9 4-4.9 5-5.9 9-9.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 2 ....... 5 ....... 3-3.9 4-4.9 5-5.9 9-6.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11. 8 12-12.9 2 ....... tt ....... 3 ....... 3-3:9 4-4.9 5-5.9 9-6.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-129 12 1 ...... 3 ....... 7 5 1 1 .... 1 ....... 3-3.9 4-4.9 5-5.9 6-6.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 1 ....... 3-3.9 4-4.9 5-5.9 6-6.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 1 ... . . 13-13.9 14-14.9 15-15.9 16-16.9 17-17.9 18-18.9 19-19.9 20-20.9 13-13.9 14-14.9 15-15.9 19-16.9 17-17.9 18-18.9 19.19.9 20-20.9 1J-13.9 14-14.9 15-15.9 16-16.9 17-17.9 18-18.9 19-19.9 20-20.9 13-13.9 14-14.9 15-15.9 19-19.9 17-17.9 18-18.9 19-19.9 20-20.9 13-13.9 14-14.9 15-15.9 19-16.9 17-17.9 18-18.9 19-19.9 20-20.9 13-13.9 14-14.9 15-15.9 19-19.9 17-17.9 18-18.9 19-19.9 20-20.9 13-13.9 14-14.9 15-15.9 19-16.9 17-17.9 18-18.9 19-19.9 20-20.9 13-13.9 14-14.9 15-15.9 19-19.9 17-17.9 18-18.9 19-19.9 20-20.9 29-26.9 37-37.9 29-28.9 37-37.9 26-26.9 37-379 26-26.9 37-37.9 26-26.9 37-37S 29-26.9 37-37* 28-29.9 37-373 r i r i r i r i r i r i r i r i i i r i r i i i i i t i r i Agua Hedionda Lagoon Fish Size Data April 1995 Species: Paralichthys eaffomieus Section Station C B3 C B4 C S1 C S2 B6 B7 S3 S7 0 B1 0 O1 Total Weight (gms) 114.1 244.15 22.3 80.3 0.15 70 13 5 0.05 50 Total Count 5 6 11 8 2 Species: Pleuronichthys ritteri Section Station C B3 Total Weight (gms) 0.1 Total Count 1 Species: Hypsopsetta guttulata Section Station C B5 C S1 2 S2 S4 S5 S9 S7 Total Weight (gms) 38 85.6 40.25 34.357.2 8.15 43.55 Total Count 1 35 17 19 1725 31 3 Species: Symphunj* ablcauda Section Station C B3 Total Weight (gms) 3 Total Count 1 Counts by Size Class (em) 0-0.9 1 1-1.91 2 1 1 2-2.9 4 2 1 3-3.9 2 1 Counts by Size Class (em) 0-0.9 1-1.9 1 2-2.9 3-3.9 4-4.9 5-5.9 6-6.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 13-13.9 14-14.9 15-15.9 16-16.9 2 .... .11. 11.. 1 . . 1 1 . . . . . . . 1211. .... ....... . '. ... 1 1 . . .... 1 . . ... 1 4-4.9 5-5.9 6-6.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 13-13.9 14-14.9 15-15.9 16-16.9 17-17.9 18-19.9 1 1 17-17.9 18-18.9 Counts by Size Class (cm) 0-0.9 1-1.9 10 2 1 15 19 1 2-2.9 9 8 4 8a 3-3.9 3 1 5 1 4 4-4.9 5-5.9 6-6.9 7-7.9 8-8.9 9-9.9 10-10.9 11-11.9 12-12.9 13-13.9 14-14.9 15-15.9 19-18.9 1 .... 3424... .. ... 1 2 1 2 ... . . . . . 232.... .. ...1 ...... . . . . . 1 ..... .1 ... 17-17.9 19-18.9 Counts by Size Class (cm) 0-0.9 1-1.9 2-2.9 3-3.9 4-4.9 5-5.9 6-6.9 7-7.9 9-8.9 9-9.9 10-10.9 11-11.9 12-12.9 13-13.9 14-14.9 15-15.9 16-16.9 1 ... . . .... 17-17.9 19-19.9 19-19.9 20-20.9 26-J0.9 37-37.9 19-19.9 20-20.9 19-19.9 20-20.9 26-29.9 37-375 26-26.9 37-37.9 19-19.9 20-20.9 26-29.9 3747.9 mi i C.1 Macro in vertebrate Density Data by Lagoon Section, Station, and Survey Agua Hedionda Lagoon Macroinvertebrate Data Density per m2 by Gear Type and Station Outer Lagoon July 1994 B1 82 Eudendrium sp. 0.004 Actiniaria 0.020 Prosthiostomum latocelis 0.004 Aphrodita armifera 0.004 Harmothoe hirsuta 0.024 Paleanotus bellis 0.004 Hesionidae 0.004 Amphiduros pacificus 0.004 Exogone lourei 0.020 Odontosyllis phosphorea 0.004 Platynereis bicanaliculata 0.004 Diopatra spp. 0.004 Leitoscoloplos pugettensis 0.004 Spiophanes missionensis 0.004 Polyophthalmus pictus 0.008 Mediomastus sp. 0.004 Ampharete labrops 0.004 Pista alata 0.012 Euchone limnicola 0.004 Oligochaeta 0.004 Tricolia sp. 0.024 Macrarenesp. 0.016 Alvania rosana 0.004 Asperiscala sp. 0.004 Crepidula sp. 0.032 Acanthina spinata 0.024 Aliacarinata 0.136 Olivella baetica 0.008 Granulina margaritula 0.004 Kurtziella plumbea 0.004 Ophiodermella incisa 0.004 Conus califomicus 0.012 Navanax inermis 0.008 Bulla gouldiana 0.032 Haminaea vesicula 0.008 Mytilus galloprovincialis 0.020 Modiolussp. 0.012 Leptopecten latiauratus 0.008 Limaria hemphilli 0.004 Laevicardium substriatum 1.456 Tellina sp. 0.004 Theora lubrica 0.004 Venerupis phillipinarum 0.008 Hiatella arctica 0.004 Anoplodactylus erectus 0.008 Anoplodactylus califomica Anoropallene palpida 0.020 Parasterope bamesi 0.004 Metamysidopsis elongata 0.004 Zeuxo sp. 0.020 Leptochelia dubia 0.112 Leptochelia cf. sp. B 0.016 Paracerceis sculpta 0.148 Neastacilla califomica 0.004 Aoroides cf. secunda 0.024 Acuminodeutopus heteruropus 0.020 Corophium spp. 0.012 Ericthonius brasiliensis . 0.024 Grandidierella japonica 0.012 Pontogeneia rostrata 0.028 Elasmopus sp. 0.008 Hyalesp. 0.016 0.003 0.010 0.003 0.003 0.010 0.003 0.003 Agua Hedionda Lagoon Macroinvertebrate Data Density per m2 by Gear Type and Station Outer Lagoon July 1994 - Continued Photis sp. 0.004 Gammaropsis thompsoni 0.036 Monoculodes hartmanae 0.004 Podocerus spp. 0.112 Stenothoe estacola 0.004 Mayerella banksia 0.008 Caprella califomica 0.004 Palaemonella holmes! 0.032 Neocrangon zacae 0.004 Pagurusspp. 0.128 Loxomynchus spp. 0.012 Cancer sp. 0.004 Cancer anthonyi 0.004 Lophopanopeus sp. 0.084 Membranipora sp. (colonial) 0.004 Microporina sp. (colonial) 0.004 Thalamoporella California (colonial) 0.004 Ophiuroidea 0.008 Amphiodia digitata 0.012 Amphipholis spp. 0.008 Amphipholis squamata 0.008 Ophiothrix spiculata 0.004 Parastichopus parvimensis 0.004 TOTAL 2.960 Number of species 80.000 0.003 0.007 0.003 0.050 10.000 Agua Hedionda Lagoon Macroinvertebrate Data Density per m2 by Gear Type and Station Middle Lagoon July 1994 B3 B4 B5 Notoacmea dipicta Crepidula sp. Crepidula onyx Alia carinata Nassarius tegula Conus califomicus Navanax inermis Bulla gouldiana Haminaea vesicula Mytilus galloprovincialis Pecten sp. Argopecten circularis Leptopecten latiauratus Laevicardium substriatum Chione undatella Leptochelia dubia Paracerceis sculpta Amphilochus neapolitanus Ampithoe spp. Acumlnodeutopus heteruropus Corophium spp. Ericthonius brasiliensis Pontogeneia rostrata Elasmopus sp. Hyale sp. Podocerus spp. Periclimenes infraspinus Hippolyte califomiensis Hemigrapsus oregonensis Lytechinus pictus TOTAL Number of species 0.004 0.004 0.044 0.052 3.000 0.008 0.016 0.048 0.028 0.020 0.004 0.064 0.004 0.004 0.004 0.004 0.188 0.008 0.024 0.012 0.220 0.008 0.052 0.576 0.064 0.008 3.052 0.304 0.088 0.004 4.812 25.000 0.016 0.012 0.012 0.012 0.016 0.004 0.004 0.200 0.004 0.008 0.108 0.024 0.004 0.424 13.000 Agua Hedionda Lagoon Macroinvertebrate Data Density per m2 by Gear Type and Station West Inner Lagoon July 1994 B6 B9 Notoacmea dipicta . 0.010 Alia carinata . 0.023 Nassarius tegula 0.008 0.067 Bulla gouldiana 0.532 0.720 Mytilus galloprovincialis 0.004 0.027 Musculista senhousi 0.008 Leptopecten latiauratus 0.004 0.020 Laevicardium substriatum . 0.113 Tellina sp. B (SCAMIT) . 0.007 Chione fluctifraga . 0.003 Chione undatella . 0.003 Cyclopoida (copepod) . 0.007 Zeuxo sp. . 0.003 Paracerceis sculpta . 0.817 Amphilochus neapolitanus . 0.037 Ampithoe spp. . 0.400 Corophium spp. . 0.323 Ericthonius brasiliensis . 0.493 Pontogeneia rostrata . 0.027 Elasmopus sp. . 0.003 Hyalesp. . 1.807 Gammaropsis thompsoni . 0.007 Podocerus spp. . 1.840 Hippolyte califomiensis 0.008 0.073 TOTAL 0.564 6.830 Number of species 6.000 23.000 East Inner Lagoon July 1994 B7 B8 Leitoscoloplos pugettensis . 0.004 Nassarius tegula 0.120 Navanax inermis 0.012 Bulla gouldiana 0.048 Musculista senhousi 0224 0.136 Tagelus affinis 0.008 Theora lubrica 0.004 Chione fluctifraga 0.008 Paracerceis sculpta 0.008 Ampithoe spp. 0.016 Grandidierella japonica 0.016 Periclimenes infraspinus 0.012 Hemigrapsus oregonensis . 0.012 TOTAL 0.476 0.152 Number of species 11.000 3.000 mi Agua Hedionda Lagoon Macroinvertebrate Data Density per m2 by Gear Type and Station Outer Lagoon April 1995 B1 Halecium sp. 0.005 Exogone lourei Lumbrineris spp. 0.005 Leltoscoloplos pugettensis 0.010 Polydora ligni 0.005 Pseudopolydora paucibranchiata Mediomastus sp. 0.005 Pistaalata 0.010 Euchone limnicola 0.015 Cerithiopsis sp. Crepidula sp. 0.005 Alia carinata 0.040 Olivella biplicata 0.010 Olivella baetica 0.035 Acteocina inculta 0.010 Melanochlamys diomedea 0.005 Bulla gouldiana 0.005 Haminaea vesicula 0.010 Musculista senhousi 0.020 Laevicardium substriatum 0.015 Mactra sp. 0.005 Mactra nasuta Ensis myrae Tellina sp. B (SCAMIT) 0.010 Theora lubrica 0.025 Transennella sp. Venerupis phillipinarum Cooperella subdiaphana Thracia curta Anoplodactylus erectus 0.015 Aoroides cf. secunda 0.005 Hippolyte califomiensis 0.010 Crangon nigromaculata 0.005 Filicrisia sp. (colonial) 0.005 Membranipora tuberculata (colonial) 0.005 Thalamoporella California (colonial) 0.005 Tricellaria sp. (colonial) 0.005 Astropecten armatus Amphiodia urtica 0.005 Leptosynapta sp. 0.005 TOTAL 0.315 Number of species 30.000 B2 0.005 0.005 0.005 0.010 0.090 0.055 0.030 0.395 0.035 0.010 0.005 0.020 0.030 0.005 0.005 0.015 0.005 0.005 0.005 0.735 18.000 mm Agua Hedionda Lagoon Macroinvertebrate Data Density per m2 by Gear Type and Station Middle Lagoon April 1995 Stylochus exiguus Stylochoplana sp. Notoplana sp. Tubulanus frenatus Eumida sp. Exogone lourei Platynereis bicanaliculata Nephtys caecoides Leitoscoloplos pugettensis Prionospio heterobranchia Diplocirrus sp. Armandia brevis Melinna oculata Pista fasciata Euchone limnicola Serpulidae Nitidiscala sp. Crepidula sp. Crepidula onyx Nassarina penicillata Alia carinata Nassarius mendicus Nassarius tegula Olivella baetica Conus califomicus Acteocina eximia Acteocina inculta Navanax inermis Bulla gouldiana Haminaea vesicula Bivalvia Nuculana taphria Musculista senhousi Laevicardium substriatum Mactra sp. Mactra nasirta Cumingia califomica Theora lubrica Venerupis phillipinarum Chione fluctifraga Chione undatella Cooperella subdiaphana Anoplodactylus erectus Diastylis sp. Monoculodes hartmanae Palaemonella holmes! Hippolyte califomiensis Crangon sp. Crangon nigricauda Crangon alaskensis Crangon nigromaculata Pagurus spp. Loxorhynchus spp. Bowerbankia sp. (colonial) Crisia sp. (colonial) Membranipora sp. (colonial) Lagenipora sp. (colonial) Amphiodia urtica Lytechinus pictus Dendraster excentricus Ascidiacea TOTAL Number of species B3 B4 B5 0.007 0.013 0.007 0.007 0.007 0.033 0.040 0.053 0.007 0.007 0.007 0.080 0.013 0.007 0.007 0.013 0.007 0.007 0.007 0.007 0.207 0.007 0.007 0.007 0.007 0.007 0.287 0.007 0.867 26.000 0.007 0.007 0.007 0.007 0.007 0.007 0.040 0.013 0.007 0.040 0.007 0.007 0.007 0.047 0.007 0.227 0.007 0.393 0.100 0.013 0.027 0.033 2.007 0.013 0.007 0.007 0.007 0.973 0.047 0.067 0.027 0.013 0.007 0.007 0.040 0.007 0.007 0.007 0.013 4.267 38.000 0.010 0.005 0.005 0.010 0.035 0.030 0.105 0.025 0.010 0.005 1.265 0.115 0.005 0.010 0.010 0.010 0.005 0.015 1.675 18.000 Agua Hedionda Lagoon Macroinvertebrate Data Density per m2 by Gear Type and Station West Inner Lagoon April 1995 Leitoscoloplos pugettensis Prionospio heterobranchia Diplocirais sp. Armandia brevis Pista alata Euchone limnicola Alia carinata Nassarius tegula Acteocina inculta Navanax inermis Bulla gouldiana Haminaea virescens Musculista senhousi Laevicardium substriatum Tellina sp. B (SCAMIT) Theora lubrica Chione undatella Anoplodactylus erectus Mysida Ampithoe spp. Grandidierella japonica Palaemonella holmes! Hippolyte califomiensis Leptosynapta sp. TOTAL Number of species B6 0.020 0.005 0.005 0.010 0.005 0.025 0.005 0.045 0.010 0.005 0.045 0.010 0.020 0.245 0.030 0.050 0.010 0.265 0.005 0.005 0.020 0.020 0.005 0.865 23.000 B9 0.010 0.010 0.010 0.010 0.010 0.020 0.060 0.010 0.010 0.020 0.170 10.000 mm mm m Agua Hedionda Lagoon Macroinvertebrate Data Density per m2 by Gear Type and Station East Inner Lagoon April 1995 Lumbrineris spp. Leitoscoloplos pugettensis Polydora ligni Pista alata Euchone limnicola Notoacmea dipicta Nassarius tegula Acteocina inculta Melanochlamys diomedea Bulla gouldiana Haminaea vesicula Bivalvia Musculista senhousi Laevicardium substriatum Mactra sp. Tellina sp. B (SCAMIT) Cumingia califomica Theora lubrica Cooperella subdiaphana Anoplodactylus erectus Mysida Paracerceis sculpta Cymothoidae Palaemonella holmesi Hippolyte califomiensis Loxorhynchus spp. Leptosynapta sp. TOTAL Number of species B7 0.040 0.080 0.010 0.170 0.060 4.740 0.430 0.010 0.260 0.160 0.020 0.010 0.030 0.140 0.030 0.030 0.200 0.010 0.010 0.030 0.010 6.480 20.000 B8 0.010 0.010 0.050 0.010 0.040 0.370 0.010 0.880 0.010 0.120 0.040 0.030 0.020 0.020 0.020 0.010 1.650 16.000 I C.2 Macroinvertebrate Total Count Data by Lagoon Section, Station, and Survey Agua Hedionda Lagoon Macroinvertebrate Data Total Counts by Gear Type and Station Outer Lagoon July 1994 B1 B2 Eudendrium sp. 1 Actiniaria 5 Prosthiostomum latocelis 1 Aphrodita armifera 1 Harmothoe hirsuta 6 Paleanotus bellis 1 Hesionidae 1 Amphiduros pacificus 1 Exogone lourei 5 Odontosyllis phosphorea 1 Platynereis bicanaliculata 1 Oiopatra spp. 1 Leitoscoloplos pugettensis 1 Spiophanes misslonensls 1 Polyophthalmus pictus 2 Mediomastus sp. 1 Ampharete labrops 1 Pista alata 3 Euchone limnicola 1 Oligochaeta 1 Tricolia sp. 6 Macrarene sp. 4 Alvania rosana 1 Asperiscala sp. 1 Crepidula sp. 8 Acanthi na splnata 6 1 Alia carinata 34 Olivella baetica 2 Granulina margaritula 1 Kurtziella plumbea 1 Ophiodermella incisa 1 Conus califomicus 3 3 Navanax inermis 2 1 Bulla gouldiana 8 Haminaea vesicula 2 Mytilus galloprovincialis 5 Modiolus sp. 3 Leptopecten latiauratus 2 1 Limaria hemphilli 1 Laevicardium substriatum 364 3 Tellina sp. 1 Theora lubrica 1 Venerupis phillipinaaim 2 Hiatella arctica 1 Anoplodactylus erectus 2 Anoplodactylus califomica . 1 Anoropallene palpida 5 Parasterope bamesi 1 Metamysidopsis elongata 1 Zeuxo sp. 5 Leptochelia dubia 28 Leptochelia cf. sp. B 4 Paracerceis sculpta 37 1 Neastacilla californica 1 Aoroides cf. secunda 6 Acuminodeutopus heteruropus 5 Corophium spp. 3 Ericthonius brasiliensis 6 Grandidierella japonica 3 Pontogeneia rostrata 7 Elasmopus sp. 2 Hyale sp. 4 Photis sp. 1 Agua Hedionda Lagoon Macroinvertebrate Data Total Counts by Gear Type and Station Outer Lagoon July 1994-Continued Gammaropsis thompsoni 9 Monoculodes hartmanae 1 Podocerus spp. 28 Stenothoe estacola 1 Mayerella banksia 2 Caprella califomica 1 Palaemonella holmesi 8 1 Neocrangon zacae 1 Pagurus spp. 32 2 Loxorhynchus spp. 3 Cancer sp. 1 Cancer anthonyi 1 Lophopanopeus sp. 21 Membranipora sp. (colonial) 1 Microporina sp. (colonial) 1 Thalamoporella California (colonial) 1 Ophiuroidea 2 Amphiodia digitata 3 Amphipholis spp. 2 Amphipholis squamata 2 Ophiothrix spiculata 1 1 Parastichopus parvimensis 1 TOTAL 740 15 Number of species 80 10 Agua Hedionda Lagoon Macroinvertebrate Data Total Counts by Gear Type and Station Middle Lagoon July 1994 B3 B4 B5 Notoacmea dipicta . 2 Crepidulasp. . 4 Crepidula onyx . . 4 Aliacarinata 1 12 3 Nassarius tegula . 7 3 Conus califomicus . 5 3 Navanax inermis . 1 Bulla gouldiana . 16 4 Haminaea vesicula . 1 Mytilus galloprovincialis . 1 Pectensp. . 1 Argopecten circularis . . 1 Leptopecten latiauratus . 1 1 Laevicardium substriatum . 47 50 Chione undatella . . 1 Leptochelia dubia . 2 Paracerceis sculpta . 6 Amphilochus neapolitanus . 3 Ampithoe spp. . 55 Acuminodeutopus heteruropus . 2 Corophium spp. . 13 Ericthonius brasiliensis . 144 Ponlogeneia rostrata . 16 Elasmopus sp. . 2 Hyalesp. 1 763 Podocerus spp. . 76 Periclimenes infraspinus . . 2 Hippolyte califomiensis 11 22 27 Hemigrapsus oregonensis . . 6 Lytechinus pictus . 1 1 TOTAL 13 1203 106 Number of species 3 25 13 Agua Hedionda Lagoon Macroinvertebrate Data Total Counts by Gear Type and Station West Inner Lagoon July 1994 B6 B9 Notoacmea dipicta . 3 Alia carinata . 7 Nassarlus tegula 2 20 Bulla gouldiana 133 216 Mytilus galloprovincialis 1 8 Musculista senhousi 2 Leptopecten latiauratus 1 6 Laevicardium substriatum . 34 Tellina sp. B (SCAMIT) . 2 Chione fluctifraga . 1 Chione undatella . 1 Cyclopoida (copepod) . 2 Zeuxo sp. . 1 Paracerceis sculpta . 245 Amphilochus neapolilanus . 11 Ampithoe spp. . 120 Corophium spp. . 97 Ericthonius brasiliensis . 148 Pontogeneia rostrata . 8 Elasmopus sp. . 1 Hyale sp. . 542 Gammaropsis thompsoni . 2 Podocerus spp. . 552 Hippolyte califomiensis 2 22 TOTAL 141 2049 Number of species 6 23 East Inner Lagoon July 1994 B7 B8 Leitoscoloplos pugettensis . 1 Nassarius tegula 30 Navanax inermis 3 Bulla gouldiana 12 Musculista senhousi 56 34 Tagelus affinis 2 Theora lubrica 1 Chione fluctifraga 2 Paracerceis sculpta 2 Ampithoe spp. 4 Grandidierella japonica 4 Periclimenes infraspinus 3 Hemigrapsus oregonensis . 3 TOTAL 119 38 Number of species 11 3 Agua Hedionda Lagoon Macroinvertebrate Data Total Counts by Gear Type and Station Outer Lagoon April 1995 B1 B2 Halecium sp. 1 Exogone lourei . 1 Lumbrineris spp. 1 Leitoscoloplos pugettensis 2 Polydora ligni 1 Pseudopolydora paucibranchiata . 1 Mediomastus sp. 1 Pista alata 2 Euchone limnicola 3 Cerithiopsis sp. . 1 Crepidula sp. 1 Alia carinata 8 2 Olivella blplicata 2 Olivella baetica 7 Acteocina inculta 2 18 Melanochlamys diomedea 1 Bulla gouldiana 1 Haminaea vesicula 2 11 Musculista senhousi 4 6 Laevicardium substriatum 3 79 Mactra sp. 1 7 Mactra nasuta . 2 Ensis myrae . 1 Tellina sp. B (SCAMIT) 2 4 Theora lubrica 5 6 Transennella sp. . 1 Venerupis phillipinarum . 1 Cooperella subdiaphana . 3 Thracia curta . 1 Anoplodactylus erectus 3 Aoroides cf. secunda 1 Hippolyte califomiensis 2 Crangon nigromaculala 1 Filicrisia sp. (colonial) 1 Membranipora tuberculata (colonial) 1 Thalamoporella California (colonial) 1 Tricellaria sp. (colonial) 1 Astropecten armatus . 1 Amphiodia urtica 1 Leptosynapta sp. 1 1 TOTAL 63 147 Number of species 30 18 Agua Hedionda Lagoon Macroinvertebrate Data Total Counts by Gear Type and Station Middle Lagoon April 1995 B3 B4 B5 Stylochus exiguus . 1 Stylochoplana sp. . 1 Notoplanasp. 1 Tubulanus frenatus . 1 Eumidasp. 2 Exogone lourei . 1 Platynereis bicanaliculata . 1 . Nephtys caecoides . 1 Leitoscoloplos pugettensis . 6 2 Prionospio heterobranchia . 2 1 Diplocirrus sp. . 1 Armandia brevis . 6 Melinna oculata . 1 Pista fasciata . . 1 Euchone limnicola . 1 2 Serpulidae 1 Nitidiscala sp. 1 1 Crepidulasp. 1 Crepidula onyx . 7 7 Nassarina penicillata . 1 Aliacarinata 5 34 6 Nassarius mendicus . 1 Nassarius tegula 6 59 21 Olivella baetica 8 Conus califomicus . 15 5 Acteocina eximia . 2 Acteocina Inculta . 4 2 Navanax inermis . 5 1 Bulla gouldiana 1 301 253 Haminaea vesicula 1 2 Bivalvia 1 1 Nuculana taphria . 1 Musculista senhousi . 1 Laevicardium substriatum 12 146 23 Mactrasp. . 7 1 Mactra nasuta 2 Cumingia califomica 1 Theora lubrica . 10 2 Venerupis phillipinarum 1 . 2 Chione fluctifraga 2 Chione undatella . 4 2 Cooperella subdiaphana . 2 Anoplodactylus erectus . . 1 Diastylissp. . 1 Monoculodes hartmanae 1 Palaemonella holmes! 1 1 Hippolyte californiensis . 6 3 Crangon sp. 1 . . Crangon nigricauda . 1 Crangon alaskensis 1 Crangon nigromaculata 31 Pagurusspp. . 1 Loxorhynchus spp. 1 Bowerbankia sp. (colonial) 1 Crisia sp. (colonial) 1 Membranipora sp. (colonial) 1 Lagenipora sp. (colonial) 1 Amphiodia urtica . 1 Lytechinus pictus . 2 Dendraster excentricus .43 Ascidiacea 1 TOTAL 130 640 335 Number of species 26 38 18 Agua Hedionda Lagoon Macroinvertebrate Data Total Counts by Gear Type and Station West Inner Lagoon April 1995 B6 B9 Leitoscoloplos pugettensis 4 1 Prionospio heterobranchia 1 Diplocirrus sp. 1 Armandia brevis 2 Pista alata 1 Euchone limnicola 5 1 Alia carinata 1 Nassarius tegula 9 Acteocina inculta 2 Navanax inermis 1 1 Bulla gouldiana 9 1 Haminaea virescens 2 Musculista senhousi 4 1 Laevicardium substriatum 49 2 Tellina sp. B (SCAMIT) 6 6 Theora lubrica 10 1 Chione undatella . 1 Anoplodactylus erectus 2 Mysida 53 Ampithoe spp. 1 Grandidierella japonica 1 Palaemonella holmesi 4 Hippolyte califomiensis 4 Leptosynapta sp. 1 2 TOTAL 173 17 Number of species 23 10 Agua Hedionda Lagoon Macroinvertebrate Data Total Counts by Gear Type and Station East Inner Lagoon April 1995 B7 B8 Lumbrineris spp. . 1 Leitoscoloplos pugettensis 4 1 Polydora ligni . 5 Pista alata . 1 Euchone limnicola 8 4 Notoacmea dipicta 1 Nassarius tegula 17 37 Acteocina inculta 6 Melanochlamys diomedea . 1 Bulla gouldiana 474 88 Haminaea vesicula 43 1 Bivalvia 1 Musculista senhousi 26 12 Laevicardium substriatum 16 4 Mactra sp. 2 Tellina sp. B (SCAMIT) 1 Cumingia califomica 3 Theora lubrica 14 3 Cooperella subdiaphana 3 Anoplodactylus erectus 3 Mysida 20 2 Paracerceis sculpta 1 Cymothoidae 1 Palaemonella holmesi . 2 Hippolyte californiensis 3 2 Loxorhynchus spp. 1 Leptosynapta sp. . 1 TOTAL 648 165 Number of species 20 16 C.3 Sediment Data for Infauna Samples Agua Hedionda Lagoon Grain Size Data Benthic Cores (Percent of Total) April 1995 Lagoon Section Outer Lagoon Outer Lagoon Middle Lagoon Middle Lagoon Middle Lagoon West Inner Lagoon West Inner Lagoon West Inner Lagoon West Inner Lagoon East Inner Lagoon East Inner Lagoon East Inner Lagoon East Inner Lagoon Sediment Type Eelgrass Bed Non-Vegetated Eelgrass Bed Bar Non-Vegetated Bar Eelgrass Bed Non-Vegetated Non-Vegetated Eelgrass Bed Non-Vegetated Bar Bar Station 1 4 5 8 9 12 14 15 18 19 22 23 25 Gravel 0.01 0.08 0.03 0.00 0.03 0.14 0.03 0.03 0.00 0.00 0.15 0.00 0.21 Sand 92.57 80.46 83.37 93.77 97.45 77.25 52.10 88.13 15.34 8.83 23.38 44.90 35.85 Silt 3.90 13.61 8.36 3.46 0.69 12.42 27.88 5.61 31.69 31.12 35.46 25.08 28.14 Clay 3.52 5.85 8.23 2.77 1.83 10.20 19.98 6.23 52.97 60.05 41.01 30.02 35.79 C.4 Infauna Density Data by Lagoon Section, Station, and Habitat II it II t 1 II f 1 II I S II I ! II II II fl II If I f 1 I I Species Agua Hedionda Lagoon Benthic Infauna Data Density per m2 Outer Lagoon Middle Lagoon Eelgrass Bed Non-Vegetated Eelgrass Bed Bar Bar 1234 5678 Non-Vegetated 9 10 Actiniaria .... Nemertea . . . 127 Lineidae .... Paranemertes californica .... Nematoda 127 6621 Polychaeta 382 637 Polynoidae .127 Tenonia priops . 127 Chloeia pinnata . . 127 Phyllodoce longipes .... Eteone spilotus .... Eumida sp. 2 .... Platynereis bicanaliculata .... Nephtys cornata franciscana .... Nephtys caecoides .... Hemipodus borealis .... Goniada littorea . . . 127 Lumbrineris spp. . 255 Lumbrineris minima . . 127 Schistomeringos rudolphi .... Scoloplos acmeceps . . . 255 Leitoscoloplos pugettensis . 127 Acmira catherinae .... Polydora spp. .... Polydora ligni .... Polydora nuchalis .... Prionospio heterobranchia . 637 Apoprionospio pygmaea . . 127 127 Spiophanes missionensis 255 1273 . 127 Rhyncospio glutea 127 127 Pseudopolydora paucibranchiata . 127 Carazziella sp. . 127 Boccardiella sp. .... Boccardiella hamata .... Diplocirrus sp. .... Armandia brevis . 127 Polyophthalmus pictus . 255 Capitellidae 127 Capitella "capitata" .... Notomastus lineatus .... Notomastus hemipodus . . 509 255 Mediomastus sp. 255 1655 891 Mediomastus ambiseta . 637 637 127 Mediomastus californiensis .... Anotomastus gordiodes . 127 Pista sp. .... Pista alata .... Euchone limnicola .... Oligochaeta .... Caecum californicum 127 764 . 509 Alia carinata .... Granulina margaritula . 255 Acteocina inculta . 382 127 255 127 4329 127 255 127 127 1146 255 1273 382 255 127 127 127 127 1910 509 764 255 1783 1146 127 127 382 3056 1655 127 127 127 127 . 127 127 8021 4074 127 127 127 1 ' * II 1 I ' i 1 i 1 I J I I I 1 I I I 1 I I I I I 1 I I I I I I I I 1 i Agua Hedionda Lagoon Benthic Infauna Data Density per m2 Outer Lagoon Middle Lagoon Eelgrass Bed Non-Vegetated Eelgrass Bed Bar Bar Non-Vegetated Species 1234 56789 10 Philine sp. A .... ...... Haminaea vesicula . 127 . ...... Bivalvia . 127 . . ...... Mytilus galloprovincialis .... ...... Musculista senhousi .... ...... Laevicardium substriatum . 1655 509 255 ..... 127 Mactra sp. . 255 . . ..... 255 Tellina sp. . 1273 382 382 255 ..... Tellina sp. B (SCAMIT) . . 127 127 127 127 .... Tagelus subteres .... ...... Tagelus affinis .... ..... 382 Cooperella subdiaphana .... ..... 127 Cryptomya californica .... . 255 .... Araneida adult .... ...... Tanystylum cf. intermedium .... 127 ..... Calanoida (copepod) 127 ... ...... Oxyurostylis sp. 255 ... ...... Paracerceis sculpta . 127 . . ...... Ampithoe spp. . 127 . . ...... Ericthonius brasiliensis . 127 . . . 382 .... Grandidierella japonica .... 127 509 .... Pontogeneia rostrata .... ...... Tethygenia opata . . . . .127 Gammaropsis thompsoni 127 Mayerella banksia . . 127 . ...... Caprella mendax .... ...... Caridea 127 ... ...... Neotrypaea californiensis 127 . 127 . . 127 . . Pinnixa spp. .... ...... Heleidae pupa .... ...... Ephydridae pupa .... ...... Phoronida 255 127 .... 127 Lichenopora sp. (colonial) .... ...... Membranipora sp. (colonial) . 127 . . ...... Ophiuroidea . . . 255 ....... Ophiuroidea .... ... 127 Leptosynapta sp. .... ...... TOTAL 2037 18462 3947 2674 11205 15406 2037 509 891 7639 Number of species 9 25 10 11 12 16 4 4 2 17 1 I \ I 1 I ! I I I ! I 1 I 1 I l Species Bar 11 , , , , f f , , , Agua Hedionda Lagoon Benthic Infauna Data Density per m2 West Inner Lagoon Bar Eelgrass Bed ------ Non-Vegetated 12 13 14 15 16 17 18 Actiniaria ..... Nemertea . 127 ... Lineidae ..... Paranemertes californica 255 .... Nematoda . 2037 ... Polychaeta ..... Polynoidae ..... Tenonia priops ..... Chloeia pinnata ..... Phyllodoce longipes . . . 127 . Eteone spi lotus ..... Eumida sp . 2 . .127 Platynereis bicanaliculata . . 127 127 . Nephtys cornata franciscana .... 127 Nephtys caecoides ..... Hemipodus borealis 127 .... Goniada littorea ..... Lumbrineris spp. . 127 . . . Lumbrineris minima ..... Schistomeringos rudolphi ..... Scoloplos acmeceps ..... Leitoscoloplos pugettensis . . 127 . 891 Acmira catherinae . 127 ... Polydora spp. . . . 127 . Polydora ligni . . 2037 2165 . Polydora nuchal is ..... Prionospio heterobranchia ..... Apoprionospio pygmaea ..... Spiophanes missionensis . 382 ... Rhyncospio glutea .... 127 Pseudopolydora paucibranchiata 127 . 255 255 . Carazziella sp. ..... Boccardiella sp. . . 127 382 . Boccardiella hamata . . 255 127 . Diplocirrus sp. .... 509 Armandia brevis . 127 255 . 127 Polyophthalmus pictus ..... Capitellidae ..... Capitella "capitata" . . 637 . 637 Notomastus lineatus ..... Notomastus hemipodus 891 382 127 . 127 Mediomastus sp. . 255 . . 382 Mediomastus ambiseta ..... Mediomastus californiensis . 127 ... Anotomastus gordiodes ..... Pista sp. ..... Pista alata .... 127 Euchone limnicola .... 255 Oligochaeta 5093 4711 1528 1273 637 Caecum californicum ..... Alia carinata ..... Granulina margaritula . . 127 .. Acteocina inculta ..... 127 127 509 382 127 1019 . . . . 255 . 127 . . . . 127 127 127 891 382 127 255 127 255 1 I • I I 1 I i I I I 1 I 1 I ! I I 1 I 1 1 II I i II II Species Bar 11 Agua Hedionda Lagoon Benthic Infauna Data Density per m2 West Inner Lagoon Bar Eelgrass Bed ------ Non-Vegetated 12 13 14 15 16 17 18 Philine sp. A Haminaea vesicula Bivalvia Mytilus galloprovincialis Musculista senhousi Laevicardium substriatum Mactra sp. Tellina sp. Tellina sp. B (SCAMIT) Tagelus subteres Tagelus affinis Cooperella subdiaphana Cryptomya californica Araneida adult Tanystylum cf . intermedium Calanoida (copepod) Oxyurostylis sp. Paracerceis sculpta Ampithoe spp. Ericthonius brasiliensis Grandidierella japonica Pontogeneia rostrata Tethygenia opata Gammaropsis thompsoni Mayerella banksia Caprella mendax Caridea Neotrypaea californiensis Pinnixa spp. Heleidae pupa Ephydridae pupa Phoronida Lichenopora sp. (colonial) Membranipora sp. (colonial) Ophiuroidea Ophiuroidea Leptosynapta sp. TOTAL Number of species 127 127 127 127 127 382 127 127 127 127 . 127 127 255 127 127 255 509 127 637 5602 764 127 509 5220 127 127 127 7130 8 9549 14 13242 18 127 10823 12 127 127 4838 16 1401 5 127 2037 8 764 127 3056 10 • • mm K i *i K i K i • i & i ii ri 11 11 • i i i 11 i i 11 i i i i Agua Hedionda Lagoon Benthic Infauna Data Density per m2 East Inner Lagoon Eelgrass Bed Non-Vegetated Bar Bar Bar Bar Species 19 20 21 22 23 24 25 26 Actiniaria ........ Nemertea ........ Lineidae ........ Paranemertes californica ........ Nematoda 127 ....... Polychaeta ........ Polynoidae ........ Tenonia priops ........ Chloeia pinnata ........ Phyllodoce longipes ........ Eteone spilotus ..... 127 Eumida sp. 2 ........ Platynereis bicanaliculata ........ Nephtys cornata franciscana ........ Nephtys caecoides ........ Hemipodus borealis ........ Goniada littorea ........ Lumbrineris spp. . . 127 ..... Lumbrineris minima . . . 127 .... Schistomeringos rudolphi 127 . . 382 .... Scoloplos acmeceps ........ Leitoscoloplos pugettensis . . 255 ..... Acmira catherinae ........ Polydora spp. ........ Polydora ligni 2801 127 509 509 637 127 Polydora nuchalis ..... 1401 . 127 Prionospio heterobranchia . 127 255 ..... Apoprionospio pygmaea . . Spiophanes missionensis . 127 127 ..... Rhyncospio glutea ........ Pseudopolydora paucibranchiata ........ Carazziella sp. ........ Boccardiella sp. . . . 127 .... Boccardiella hamata ........ Diplocirrus sp. ........ Armandia brevis . . 127 . . . . Polyophthalmus pictus ........ Capitellidae ........ Capitella "capitata" . 255 127 891 891 1655 Notomastus lineatus ........ Notomastus hemipodus ........ Mediomastus sp. ........ Mediomastus ambiseta ........ Mediomastus californiensis ........ Anotomastus gordiodes ........ Pista sp. ........ Pista alata ........ Euchone limnicola . . 127 ..... Oligochaeta ........ Caecum californicum ........ Alia carinata ........ Granulina margaritula ........ Acteocina inculta . 127 637 3183 127 127 • llllilllilll I 1 f 1 I 1 I ! I 1 I I I 1 I 1 1 1 I i I I I 1 Species Agua Hedionda Lagoon Benthic Infauna Data Density per m2 East Inner Lagoon Eelgrass Bed Non-Vegetated Bar Bar Bar 19 20 21 22 23 24 25 Bar 26 Philine sp. A Haminaea vesicula Bivalvia Mytilus galloprovincialis Musculista senhousi Laevicardium substriatum Mactra sp. Tellina sp. Tellina sp. B (SCAMIT) Tagelus subteres Tagelus affinis Cooperella subdiaphana Cryptomya californica Araneida adult Tanystylum cf. intermedium Calanoida (copepod) Oxyurostylis sp. Paracerceis sculpta Ampithoe spp. Ericthonius brasiliensis Grandidierella japonica Pontogeneia rostrata Tethygenia opata Gammaropsis thompsoni Mayerella banksia Caprella mendax Caridea Neotrypaea californiensis Pinnixa spp. Heleidae pupa Ephydridae pupa Phoronida Lichenopora sp. (colonial) Membranipora sp. (colonial) Ophiuroidea Ophiuroidea Leptosynapta sp. TOTAL Number of species 127 255 127 127 255 127 127 1273 127 127 127 127 127 127 127 4584 6 891 6 127 127 3056 14 5220 6 2674 4584 6 3692 7 382 382 2 3 m m C.5 Biomass of Taxonomic Groups of Infauna by Lagoon Section, Station, and Habitat i ii ii ii it ii ii ii ii ii ii ti ir •• i ii ii ti Taxonomic Group Agua Hedionda Lagoon Benthic Infauna Data Biomass (gm/m2) Outer Lagoon Middle Lagoon Eelgrass Bed Non-Vegetated Eelgrass Bed Bar Bar 1234 5678 Non-Vegetated 9 10 Crustaceans Insects Mollusks Minor Phyla Polychaetes TOTAL 0. 0. 0. 0. 13. .255 .000 .127 .127 .751 0. 0. 17. 9. 39. .382 ,000 ,189 ,167 ,343 0 0 6, 0 15. .127 .000 .366 .127 .152 0. 0. 12. 0. 25. .000 .000 .096 .764 .083 14.260 66.081 21.772 37.943 0. 0. 23. 7. 8. 39, ,382 ,000 .046 ,257 ,913 ,598 76. 0. 0. 0. 16. 93. ,649 .000 ,255 ,127 ,425 .456 0. 0. 0. 0. 1. 1. ,000 ,000 000 ,127 783 ,910 0. 0. 0. 0. 0. 0. .000 .000 .127 .509 ,255 ,891 0. 0. 0. 0. 0. 0. .000 ,000 ,000 ,000 ,764 ,764 0. 0, 21, 1, 18. 42, .000 .000 .900 .528 .971 .399 II 1 t 1 K 1 ft • mm 11 ii vi 11 11 ii ii ii 11 Taxonomic Group Bar 11 Agua Hedionda Lagoon Benthic Infauna Data Biomass (gm/m2) West Inner Lagoon Bar Eelgrass Bed Non-Vegetated 12 13 14 15 16 17 18 Crustaceans Insects Mollusks Minor Phyla Polychaetes TOTAL 37.561 0.000 0.127 1.146 36.287 75.121 2.419 0.000 5.984 0.637 11.332 20.372 19. 0, 5, 0. 27. 52 .353 .000 .475 .000 .247 .076 18. 0. 67, 0. 7, 93. .080 ,000 .864 ,127 .257 .329 0 0 12 2 26 40 .000 .000 .223 .292 .356 .871 0.000 0.000 0.000 4.329 4.329 8.658 0. 0, 0. 0, 5, 6 ,127 .000 .637 .127 .220 .112 0.000 0.000 10.568 3.183 5.220 18.971 § i ii r i t i vi ti r i ii r i ri t i 11 • i ft i ri r i i i i i i i Agua Hedionda Lagoon Benthic Infauna Data Biomass (gm/m2) East Inner Lagoon Eelgrass Bed Non-Vegetated Bar Bar Bar Bar Taxonomic Group 19 20 21 22 23 24 25 26 Crustaceans 1.146 0.127 0.127 0.000 0.127 0.000 0.000 0.000 Insects 0.000 0.000 0.000 0.000 0.000 0.127 0.127 0.127 Mollusks 1.528 1.146 2.801 22.154 5.984 8.021 3.183 0.000 Minor Phyla 0.127 0.127 2.037 0.255 4.456 0.255 0.000 0.000 Polychaetes 2.037 1.146 4.456 8.403 3.438 15.406 0.000 0.127 TOTAL 4.838 2.546 9.422 30.812 14.006 23.810 3.310 0.255